USP2 is associated with mitochondrial respiratory activity and cell viability in endometrial stromal cells in endometriosis

In: BMC Women's Health · 2026 · doi:10.1186/s12905-026-04781-9 · W7196979208
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US2 was more highly expressed in ectopic endometrial stromal cells from women with endometriosis, where its manipulation altered mitochondrial respiratory activity and cell proliferation.

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This study investigated the role of ubiquitin-specific protease 2 (USP2) in mitochondrial function and proliferation within endometrial stromal cells isolated from patients with ovarian endometriosis. The researchers compared paired eutopic and ectopic tissues, finding that USP2 expression was significantly higher in ectopic lesions, which also exhibited enhanced respiratory activity and cell viability. Experimental manipulation of USP2 levels demonstrated that it directly modulates oxygen consumption rates, ATP production, and reactive oxygen species levels by regulating mitochondrial complex IV subunits. This paper is centrally about endometriosis — specifically examining the molecular mechanisms driving the proliferative and metabolic phenotype of ectopic endometrial stromal cells.

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Abstract

Mitochondrial dysfunction has been implicated in the pathophysiology of endometriosis, but the role of ubiquitin-specific protease 2 (USP2) remains unclear. This study aimed to determine whether USP2 modulates mitochondrial respiratory activity and proliferation in ectopic endometrial stromal cells (ESCs). Twenty women with ovarian endometriosis who underwent laparoscopic surgery at the Department of Obstetrics and Gynecology, Xuzhou Central Hospital, between April 2024 and April 2025 were enrolled. Paired eutopic endometrial and ectopic endometriotic tissues were collected from each participant. Primary ESCs were isolated and cultured, and USP2 expression was downregulated or upregulated by lentiviral transduction. USP2 mRNA levels were quantified by reverse-transcription quantitative polymerase chain reaction, whereas the protein levels of USP2 and the mitochondrial complex IV subunits COX5B and COX6C were assessed by western blotting. Cell viability and proliferative activity were evaluated using the Cell Counting Kit-8 assay. Intracellular reactive oxygen species (ROS) levels were measured by flow cytometry, and the oxygen consumption rate (OCR) was determined using a Seahorse extracellular flux analyzer. Intracellular ATP content and mitochondrial complex IV enzymatic activity were measured using commercial assay kits. Compared with paired eutopic ESCs, ectopic ESCs exhibited greater proliferative activity, lower intracellular ROS levels, a higher OCR, increased intracellular ATP content, and elevated mitochondrial complex IV activity (all P < 0.01). In ectopic ESCs, USP2 knockdown reduced proliferative activity ( P < 0.001), increased intracellular ROS levels ( P < 0.01), and decreased OCR, intracellular ATP content, mitochondrial complex IV activity, and the protein levels of the complex IV subunits COX5B and COX6C (all P < 0.01). Conversely, USP2 overexpression in eutopic ESCs produced reciprocal changes, including increased proliferative activity, reduced intracellular ROS levels, and higher OCR, ATP content, complex IV activity, and COX5B and COX6C protein levels (all P < 0.01). Treatment with carbonyl cyanide 4-(trifluoromethoxy) phenylhydrazone (FCCP), a mitochondrial uncoupler, attenuated the cellular and mitochondrial changes associated with USP2 overexpression. USP2 was more highly expressed in ectopic than in paired eutopic endometrial tissues. Its manipulation altered ectopic ESC proliferation and mitochondrial respiratory parameters, suggesting a potential role for USP2 in the proliferative and mitochondrial phenotype of ectopic ESCs.
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Abstract

Background Mitochondrial dysfunction has been implicated in the pathophysiology of endometriosis, but the role of ubiquitin-specific protease 2 (USP2) remains unclear. This study aimed to determine whether USP2 modulates mitochondrial respiratory activity and proliferation in ectopic endometrial stromal cells (ESCs).

Methods

Twenty women with ovarian endometriosis who underwent laparoscopic surgery at the Department of Obstetrics and Gynecology, Xuzhou Central Hospital, between April 2024 and April 2025 were enrolled. Paired eutopic endometrial and ectopic endometriotic tissues were collected from each participant. Primary ESCs were isolated and cultured, and USP2 expression was downregulated or upregulated by lentiviral transduction. USP2 mRNA levels were quantified by reverse-transcription quantitative polymerase chain reaction, whereas the protein levels of USP2 and the mitochondrial complex IV subunits COX5B and COX6C were assessed by western blotting. Cell viability and proliferative activity were evaluated using the Cell Counting Kit-8 assay. Intracellular reactive oxygen species (ROS) levels were measured by flow cytometry, and the oxygen consumption rate (OCR) was determined using a Seahorse extracellular flux analyzer. Intracellular ATP content and mitochondrial complex IV enzymatic activity were measured using commercial assay kits.

Results

Compared with paired eutopic ESCs, ectopic ESCs exhibited greater proliferative activity, lower intracellular ROS levels, a higher OCR, increased intracellular ATP content, and elevated mitochondrial complex IV activity (all P < 0.01). In ectopic ESCs, USP2 knockdown reduced proliferative activity (P < 0.001), increased intracellular ROS levels (P < 0.01), and decreased OCR, intracellular ATP content, mitochondrial complex IV activity, and the protein levels of the complex IV subunits COX5B and COX6C (all P < 0.01). Conversely, USP2 overexpression in eutopic ESCs produced reciprocal changes, including increased proliferative activity, reduced intracellular ROS levels, and higher OCR, ATP content, complex IV activity, and COX5B and COX6C protein levels (all P < 0.01). Treatment with carbonyl cyanide 4-(trifluoromethoxy) phenylhydrazone (FCCP), a mitochondrial uncoupler, attenuated the cellular and mitochondrial changes associated with USP2 overexpression.

Conclusions

USP2 was more highly expressed in ectopic than in paired eutopic endometrial tissues. Its manipulation altered ectopic ESC proliferation and mitochondrial respiratory parameters, suggesting a potential role for USP2 in the proliferative and mitochondrial phenotype of ectopic ESCs. Abbreviations - USP2: - Ubiquitin-specific protease 2 - OXPHOS: - Oxidative phosphorylation - ATP: - Adenosine triphosphate - HRP: - Horseradish peroxidase - ESCs: - Endometrial stromal cells - FCCP: - Carbonyl cyanide 4-(trifluoromethoxy)phenylhydrazone - OCR: - Oxygen consumption rate

Acknowledgements

Not applicable. Disclosures None of the authors has any commercial associations or financial disclosures that might pose or create a conflict of interest with information presented in this article. No external funding was received. Funding Not applicable. Author information Authors and Affiliations Corresponding author Ethics declarations Ethics approval and consent to participate The study was conducted in accordance with the Declaration of Helsinki and was approved by the Ethics Committee of Xuzhou Central Hospital (Approval No. XZXY-LJ-20240401-018). Written informed consent was obtained from all participants before tissue collection. Consent for publication Not applicable. Competing interests The authors declare no competing interests. Additional information Publisher’s Note Springer Nature remains neutral with regard to jurisdictional claims in published maps and institutional affiliations. Rights and permissions Open Access This article is licensed under a Creative Commons Attribution-NonCommercial-NoDerivatives 4.0 International License, which permits any non-commercial use, sharing, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if you modified the licensed material. You do not have permission under this licence to share adapted material derived from this article or parts of it. The images or other third party material in this article are included in the article’s Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by-nc-nd/4.0/. About this article Cite this article Sun, Q., Cao, Y., Gu, J. et al. USP2 is associated with mitochondrial respiratory activity and cell viability in endometrial stromal cells in endometriosis. BMC Women's Health (2026). https://doi.org/10.1186/s12905-026-04781-9 Received: Accepted: Published: DOI: https://doi.org/10.1186/s12905-026-04781-9

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