Cloacal scent gland secretions are effective in a breeding season but not sex-specific in Japanese rat snake (Elaphe climacophora) (Chordata, Reptilia, Squamata, Colubridae)

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Preprint ARPHA Preprints https://doi.org/10.3897/arphapreprints.e196454 (21 Apr 2026) https://doi.org/10.3897/arphapreprints.e196454 (21 Apr 2026) Submitted to Herpetozoa Other versions: - Preprint InfoPreprint Info - CiteCite - MetricsMetrics - CommentComment - RelatedRelated - CitedCited ARPHA Preprints doi: 10.3897/arphapreprints.e196454 First posted 21 Apr 2026 Authors Hiroki Akimoto - Corresponding author Graduate School of Advanced Science and Engineering, Waseda University, Tokyo, Japan Faculty of Education and Integrated Arts and Sciences, Waseda University, Tokyo, Japan Conflict of interest The authors have declared that no competing interests exist. Supporting agencies JSPS - Japan Society for the Promotion of Science Waseda Research Institute for Science and Engineering This is an open access preprint distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.

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