{"paper_id":"18872c06-e05d-4bc0-b0f1-e64967b3f79a","body_text":"Preprint\nARPHA Preprints\nhttps://doi.org/10.3897/arphapreprints.e196454 (21 Apr 2026)\nhttps://doi.org/10.3897/arphapreprints.e196454 (21 Apr 2026)\nSubmitted to Herpetozoa\nOther versions:\n- Preprint InfoPreprint Info\n- CiteCite\n- MetricsMetrics\n- CommentComment\n- RelatedRelated\n- CitedCited\nARPHA Preprints\ndoi:\n10.3897/arphapreprints.e196454\nFirst posted\n21 Apr 2026\nAuthors\nHiroki Akimoto\n- Corresponding author\nGraduate School of Advanced Science and Engineering, Waseda University, Tokyo, Japan\nFaculty of Education and Integrated Arts and Sciences, Waseda University, Tokyo, Japan\nConflict of interest\nThe authors have declared that no competing interests exist.\nSupporting agencies\nJSPS - Japan Society for the Promotion of Science\nWaseda Research Institute for Science and Engineering\nThis is an open access preprint distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.","source_license":"CC-BY-4.0","license_restricted":false}