Plio-Pleistocene hydrothermal events of the Baza Basin (Betic Cordillera, SE Spain) and their paleoecological implications

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These sediments were deposited in fluvio-lacustrine environments and preserve huge assemblages of large mammals, including the oldest evidence of hominin presence in Western Europe at the late Early Pleistocene sites of Barranco León and Fuente Nueva-3, dated to ∼1.4 Ma. Since latest Miocene times, the basin was subject to intense tectonic activity, showing sedimentary deposits originated from ancient hot springs. These hot springs were developed in a spatio-temporal network related to tectonic fractures, showing a higher concentration in the Orce sub-basin during Late Pliocene to Holocene times. Compositional analyses of cherts and other facies associated with these hot springs show geochemical and mineralogical markers linked to hydrothermal scenarios. The contribution of hot springs in the Baza Basin resulted in rich and productive biotopes, which supported a high diversity and biomass of terrestrial mammals. Similar geological, hydrochemical and ecological environments are found along hotspot lines in East Africa, the Levantine Corridor and the circum-Mediterranean realm. We hypothesize here that the presence of these hot springs corridors favored the dispersal of hominins and other large mammals during the Plio-Pleistocene, which led to the first hominin arrival in Europe. Figures Figure 1 Figure 2 Figure 3 Figure 4 Figure 5 Figure 6 Figure 7 Figure 8 Figure 9 1 Introduction and background The Baza Basin lies in the northeastern sector of the Guadix-Baza Depression (Granada, SE Spain; Fig. 1 A), which extends over an area of ~ 4000 km 2 (Palmqvist et al., 2022a ) and is surrounded by the highest reliefs of the Alpine Betic cordillera. This inland basin preserves a unique Plio-Pleistocene sedimentary record of high preservational completeness, which is composed of fluvio-lacustrine deposits that preserve many paleontological and archaeological localities distributed throughout the whole sedimentary depression, and especially in the Baza Basin (Martínez-Navarro, 1991 ; Palmqvist et al., 1996 , 2011 , 2022a , 2022b , 2023 ; Arribas and Palmqvist, 1998 ; Palmqvist and Arribas, 2001 ; Espigares et al., 2013 , 2019 ; Toro-Moyano et al., 2013 ; Rodríguez-Gómez et al., 2016 , 2017 ; Ros-Montoya et al., 2017 , 2018 ; Martínez-Navarro et al., 2018 ; Table 1 ). The presence of Plio-Pleistocene hydrothermal events in the Baza Basin and their paleoecological consequences has been documented since the end of the 20th century by means of sedimentological, tectonic, and mineralogical data (e.g., Sebastián Pardo, 1979 ; García-Aguilar, 1997 ; García-Aguilar et al., 2014 , 2015 ; Sánchez-Roa et al., 2016 ). 1.1 Geological and paleontological setting According to the classification of Miall ( 1984 ), the Baza Basin is an intramountainous basin with tectonic subsidence and continental sedimentation, features inherited from an ancient marine basin formed during the Late Miocene in the interior of the Betic mountain range (Sanz de Galdeano and Vera, 1992; Platt et al., 2013 ). The transition from a marine to a continental sedimentary regime took place 7.5–7.0 Ma ago, being the result of major regional tectonic movements that caused the isostatic uplifting of the basin (average uplift rate of ~ 200 m/Ma, as estimated from Late Neogene coastal marine conglomerates and coral reefs; Braga et al., 2003 ), which resulted in its disconnection from the Mediterranean Sea (Soria, 1993 ; García-Aguilar, 1997 ; Playá et al., 2000; Guerra-Merchán et al., 2014 ). The Baza Basin is bounded to the north by reliefs of Mesozoic age, mostly carbonates (Thetys type), which belong to the External Zones of the Betic Cordillera. To the south, it is surrounded by reliefs composed of sedimentary and metamorphic rocks of Paleozoic-Triassic age, which belong to the Internal Zones. To the west, it is connected with the Neogene Guadix Basin and to the southeast with the Neogene Almanzora Corridor Basin (Fig. 1 B). The Plio-Pleistocene stratigraphic model of the Baza Basin shows three major stages (Fig. 1 C), which are represented by tectono-sedimentary units (TSU) according to Megías ( 1982 ) (see also Guerra-Merchán, 1992 ; Soria, 1993 ; García-Aguilar, 1997 ; García-Aguilar and Martín, 2000 ; García-Aguilar and Palmqvist, 2011 ). The three TSUs are bounded by sedimentary hiatuses and/or unconformities that represent stages in which the basin was subject to intense tectonic activity, showing little or no sedimentation. These units have been dated using biochronological (e.g., Alberdi and Bonadonna, 1989 ; Martínez-Navarro, 1991 ; Sesé, 1991 ; Guerra-Merchán, 1992 ; Oms et al., 2000a ; Sesé, 2006 ; Gibert and Gibert, 2003 ; Ros-Montoya, 2010 ; Maldonado-Garrido et al., 2017 ) and paleomagnetic data (e.g., Garcés, 1994 ; Oms et al., 1994 , 1996 , 1999 , 2000b ; Gibert et al., 2006 , 2007 ; Scott et al., 2007 ; Oms et al., 2011 ; Álvarez et al., 2015 ; Álvarez-Posada et al., 2017 ). The first TSU, which is of Pliocene age, shows a sediment thickness of ~ 150 m. It is characterized by the alternation of marls/calcilutites/lutites and limestones deposited in lacustrine systems, which change towards the edges of the basin to sands and conglomerates of alluvial origin. Forest ecosystems were formed on the margins of the lake as inferred for the Ruscinian (~ 4.5 Ma) Baza-1 paleontological site (Ros-Montoya et al., 2017 ). Tectonic activity during this stage induced the formation of many soft-sediment deformation structures (i.e., paleo-seismites), which are mainly located at the marginal sites of a paleo-lake (Alfaro et al., 1997 ) and possibly resulted in deposits with hydrothermal affinities (García-Aguilar et al., 2014 , 2015 ). Climate was warm and humid during this stage, which relates to an environment typical of modern subtropical or rainy tropical climates (Foucault and Mélières, 2000 ; Haywood and Valdés, 2004 ). The second TSU has a latest Pliocene to Middle Pleistocene age and is the most complex, both lithologically and tectonically. During this stage, a great paleogeographic change took place in the Baza Basin, which involved an ~ E-W expansion linked to an NNW-SSE fracture system. This in turn resulted in intense subsidence and the deposition of sediments with a minimum thickness of ~ 280 m in the central sector of the basin. However, geophysical data (Sanz de Galdeano et al., 2007) indicate that sedimentary thickness may be much greater than the one visible. These sediments, contributed by fluvial systems from the adjacent Guadix Basin, filled a large lake of marly-evaporitic sedimentation with an outer rim of detrital inputs (Vera, 1970 ; García-Aguilar and Martín, 2000 ; García-Aguilar and Palmqvist, 2011 ). Tectonic activity during this stage resulted in synsedimentary normal faults, tilting, local unconformities, and seismites (Vera et al., 1984 ; Alfaro et al., 1997 , 2000 ). The lower section of this unit shows selenitic and microcrystalline gypsum levels of centimeter thickness, which alternate with layers of light-colored marls towards the basin center or with sandstones towards the edges (García-Aguilar et al., 2013 ). Climate at this stage was semi-arid, with cyclic oscillations within the Bond frequency timing band (Hagelberg et al., 1994 ; Bond et al., 1997 ; García-Aguilar et al., 2013 ), which generated alternating periods of inundation and desiccation of the lake linked to the deposition of marls or sandstones and gypsum, respectively. The upper section of this TSU outcrops mainly in the Orce area, a sub-basin in the NE sector of the Baza Basin, which has an extent of ~ 170 km 2 (Figs. 1 B–C) and shows extensive outcrops ~ 35 m thick that correspond to the Early to Middle Pleistocene. The Orce sub-basin preserves a unique fossil record of terrestrial vertebrates, including two archaeological sites dated to ~ 1.4 Ma (see review in Palmqvist et al., 2016 ) that mark the earliest presence of hominins in Western Europe: Barranco León (BL) and Fuente Nueva-3 (FN3). BL and FN3 preserve huge assemblages of skeletal remains of large mammals and Oldowan tools, which are linked through the presence of anthropogenic cut and percussion marks, as well as a human deciduous tooth in the case of BL, which is at this moment the oldest hominin fossil found in Western Europe (Martínez-Navarro et al., 1997 , 2014 ; Oms et al., 2000a , 2000b ; Espigares et al., 2013 , 2019 ; Toro-Moyano et al., 2013 ; Titton et al., 2021 ; Yravedra et al., 2021 , 2022 ; Palmqvist et al., 2023 ). Moreover, the Venta Micena site (VM, 1.6–1.5 Ma), which is slightly older than BL/FN3 and has been interpreted as a denning area of the giant hyena Pachycrocuta brevirostris (Arribas and Palmqvist, 1998 ; Palmqvist and Arribas, 2001 ; Palmqvist et al., 2022b ), provides an excellent record of the large mammal community that preceded the hominin dispersal in the basin (Martínez-Navarro, 1991 ; Mendoza et al., 2005 ; Rodríguez-Gómez et al., 2017 ). The varied lithological composition of the Orce area, with alternations of fluvial red detrital facies and lacustrine carbonate-marly facies that locally show gypsum deposits (restricted to the center of the lacustrine system in this area) and cherts, points to a paleogeographic model of fluvio-lacustrine sedimentation highly variable in time and space. This resulted in open plains with forest patches in the emerged areas surrounding the fluvio-lacustrine system (Mendoza et al., 2005 ; Scott et al., 2007 ; García-Aguilar and Palmqvist, 2011 ; Palmqvist et al., 2022a ). The third TSU of the Baza Basin has a Middle Pleistocene to Late Pleistocene age, showing a maximum thickness of 70 m. During this stage, the activity of major faults (Baza and Galera) resulted in some tectonic blocks that developed an effective subsidence, which allowed a continuous sedimentation. This TSU is composed of conglomerates and sands towards the basin edges, which change to lutites, marls and marly limestones deposited in shallow lacustrine environments towards the basin center. In the upper part of this unit and at the basin edges, alluvial facies (mainly red conglomerates and sands) prograde over lacustrine facies. This TSU is represented in several sectors of the basin perimeter (Figs. 1 B–C) and the most significant paleontological sites are Huéscar-1 and Cúllar-Baza-1 (Ruiz Bustos, 1976; Peña et al., 1977 ; Mazo et al., 1985 ; Alberdi and Bonadonna, 1989 ; Agustí et al., 2000 ; Torrente Casado, 2010 ; Ros-Montoya et al., 2018 ). Finally, during the late Middle Pleistocene (~ 400 ka: García-Tortosa et al., 2008a , b ) or in Late Pleistocene times (100–17 ka: Viseras and Fernández, 1992 ; Calvache and Viseras, 1997 ; 42 ka: Azañón et al., 2006 ), the hydrographic network of the sedimentary depression was captured by the Guadiana Menor River, a tributary of the Guadalquivir River. This resulted in the transit from an endorheic regime to an exorheic one, leading to a stage in which erosion dominated over sedimentation. As a result, a badlands landscape predominates now in the region, linked to a mean annual precipitation (MAP) of ~ 300 mm in the innermost part of the sedimentary depression ( https://es.climate-data.org ). However, climate was more humid during the late Early Pleistocene, with an estimated MAP of ~ 700 mm in BL/FN3 and ~ 850 mm in VM (Palmqvist et al., 2022c ). 1.2 Hot springs currently active in the Baza Basin Tectonic dynamism resulted in hot springs, several of which are currently active in the Baza Basin and other nearby areas (Cruz-Sanjulián et al., 1972 ; Cruz-Sanjulián and García-Rosell, 1972 , 1975 ; Diputación de Granada-ITGE, 1990 ; Figs. 1 A–B, and Table 2 ). There are twelve hot springs in the Baza Basin and four in the Guadix Basin, all of them associated with regional faults. In the case of the hot springs of the Baza Basin, their water output temperatures range between 18º C and 22º C, except in the case of the ‘Baños de Zújar’ spring, with 38º C. The hydrochemistry of the low temperature springs shows a predominance of calcium bicarbonate. In contrast, the high temperature springs mainly show silica, chlorides, and sulphates. This correspondence between high water output temperatures and siliceous-sulphate-chloride hydrochemistry appears also in the > 30º C hot springs of the Guadix Basin. The current climatic scenario of the Baza Basin shows an average annual temperature of 13.5º C (13.8º C in the Baza station and 13.3º C in the Orce station; https://es.climate-data.org ), with strong annual oscillations. Specifically, average temperature ranges between 5º C and 9º C from November to March, averaging ~ 25º C during summer. Likewise, the historical record of annual rainfall averages 335 mm at the Baza station and 366 mm at the Orce station ( https://es.climate-data.org ). In any case, the average water output temperature of the hot springs in the Baza Basin is 21º C, which is 7.5º C higher than the average annual atmospheric temperature in the basin. These data indicate that the water output temperatures of the springs are slightly lower than the average temperatures in summer, but much higher than in autumn and winter. This underlines the thermal character of these hot springs, whose flow rate varies between 1 and 400 l/s. 1.3 Tectonic dynamism From a tectonic point of view, the Baza Basin lies in the interior of the alpine-type Betic mountain range, showing three main fracture systems, which are NE-SW, NNW-SSE and NNE-SSW oriented (Sanz de Galdeano, 1983, 1990, 2008 ; Sanz de Galdeano and Vera, 1991, 1992; Sanz de Galdeano et al., 2007, 2012). These fractures played an important role in the tectono-sedimentary evolution of the Betic mountain range and generated a deep step-like geometry, which is indicative of epicortical extension processes (Sanz de Galdeano and Vera, 1992; Platt et al., 2013 ). Sanz de Galdeano et al. (2007) established two depocenters in the Baza Basin, induced by NNW-SSE and N70ºE faults that resulted in a regional ENE extension, as deduced by gravity maps of regional, residual and Bouguer anomalies: -150 mGal in Benamaurel (central zone of Baza Basin) and − 16 mGal at the north of the Orce village (central zone of Orce sub-basin). Considering basement densities of 2.67 g/cm 3 and 2.32 g/cm 3 for the sedimentary infillings of both sectors, respectively (Sanz de Galdeano et al., 2007), these result in basement depths of ~ 1300 m and ~ 500 m, respectively. In this sense, Sanz de Galdeano ( 2008 ) highlighted the Cadiz-Alicante N70ºE fault as the boundary of the inferred depocenters in the central Baza and Orce-North sectors. Likewise, Sanz de Galdeano et al. (2012, 2020) described using geological criteria the main active fractures present in the Baza Basin, which have most likely induced the historical earthquakes recorded in this region: the Galera fault (23 km long and > 5 km deep), the Baza fault (37 km long and > 10 km deep), the Botardo-Alfahuara fault (22 km long and > 5 km deep), the Benamaurel fault (19 km long and > 10 km deep), and the Cúllar faults (North and West, ~ 9 km long and 5–10 km deep, deduced in this case only by seismic criteria). Seismic phenomena were frequent during the Plio-Pleistocene, as evidenced by the presence of paleo-seismites (Alfaro et al., 1997 , 2000 , 2010 ; Gibert et al., 2005 ). Therefore, considering a standard value of the upper crustal geothermal gradient of 30º C/km (Harrison et al., 1990 ; Gupta and Roy, 2007 ), the depth calculated for these faults and the possible hydrothermal phenomena associated with them, we estimate temperatures of at least 150º–300º C in their deepest zones. 1.4 Objectives The main goal of this study is to establish a causal link between the presence of hydrothermal upwelling-related deposits in the continental Plio-Pleistocene infillings of the Baza Basin, their tectono-sedimentary origin, and their paleoecological consequences. For doing so, we have: (i) reviewed and documented the tectono-sedimentary context of the Baza Basin; (ii) performed a detailed field study, sampling selected stratigraphic sections; (iii) studied the sedimentology of these sections, analyzing the geochemical and mineralogical composition of the samples; and (iv) performed both local and high scale (Africa and Eurasia) correlations of the relationship between tectonic activity and thermal springs, on the one hand, and the presence of archaeological and paleontological sites, on the other. 2 Materials and Methods We have reviewed and documented the tectono-sedimentary context of the main archaeological and paleontological sites of the Baza Basin, as well as the sedimentary and compositional models of many modern and ancient thermal springs worldwide distributed together with the paleoanthropological sites linked to these springs. Field work in the Baza Basin consisted of geological mapping and observations at a mesoscopic scale of lithologies and their sedimentary structures, as well as of the main faults affecting the sedimentary infillings of the basin. Detailed stratigraphic profiles were obtained and the different sedimentary facies present in them were sampled (Fig. 2 ). Extensive sediment sampling was carried out along the stratigraphic record surveyed (Figs. 1 B–C, Fig. 2 ) to perform mineral and chemical analyses for identifying those facies potentially associated with a hydrothermal origin. In this study, we have analyzed 56 samples located throughout the Baza Basin, which correspond to eight types of lithologies: calcilutites, lutites, dark lutites, marls, travertines, cherts, gypsum, and native sulfur. The samples were analyzed geochemically by means of X-ray fluorescence (XRF). In addition, 14 of these samples were selected for analyzing their mineralogy by means of X-ray diffraction (XRD). All measurements were carried out at the Central Services of Support to Research (‘Servicios Centrales de Apoyo a la Investigación’, SCAI) of the University of Málaga (UMA). 2.1 Elemental analysis Chemical analyses of the abundance of major and some trace elements were carried out using a Thermo Fisher ADVANT'XP wavelength dispersive atomic fluorescence spectrometer (XRF). This equipment consists of an autosampler with capacity for a total of 161 samples, a Rh tube and a goniometer with three collimators and four crystals that allow measuring from F to U. The measurements were obtained in a He atmosphere with the sample in bead. The results were analyzed with Thermofisher's UniQuant software, which allows the semi-quantification of unknown materials without the need of standards or reference materials. Relative errors in the results range between 0.3% and 1.5% in the case of the elements with higher proportions, and up to 5% in the case of those elements with lower proportions. In order to evaluate the influence of hydrothermal activity, the ratios (Fe + Mn)/Ti and Al/(Al + Fe + Mn) were calculated (Boström et al., 1973 ; Murray, 1994 ; Madondo et al., 2021 ; Zhang et al., 2023 ). Moreover, an Al-Fe-Mn ternary diagram was also used to differentiate between hydrothermal, detrital or biogenic origins (Adachi et al., 1986 ). 2.2 Mineralogical analysis Fourteen samples, selected a priori under a hydrothermal context, were analyzed by X-ray Diffraction (XRD) based on the XRF data. The measurements were obtained with an X'Pert PRO MPD diffractometer from PANalytical and measured in high resolution conditions. The X-ray tube has a Cu anode and the diffractometer in the incident beam consists of a Johansson-type primary monochromator with a Ge (111) crystal, which provides a strictly monochromatic Cu K α1 radiation. Random powder X-ray diffraction (XRD) of bulk-rocks was used for the identification of mineral phases, measuring the diffraction angles and the intensities of the diffracted beams, which allows to compare the resulting diffraction pattern with a reference database. The comparison was carried out with PDF-4 + 2021 (Gates-Rector and Blanton, 2019 ). When the acquired data were of sufficient quality (i.e., measured with a wide range of angles, good resolution and high intensity/noise ratio), they were used not only to identify mineral phases but also to quantify them. To determine the proportion of phyllosilicate in the global samples, the phyllosilicates (4.46 Å)/quartz (3.343Å) peak-area ratio was estimated from XRD patterns of randomly oriented powder. D-spacing and the full width at half-high maximum (FWHM) were determined on the (101) reflection peak of the opal specimen in order to establish the type of opal polymorphs (Elzea et al., 1994 ). 3 Results Almost one hundred archaeological and paleontological sites with vertebrate remains dated between the Early Pliocene (Ruscinian) and the late Early Pleistocene (Galerian) have been described in the Baza Basin (e.g., Casas et al., 1975 ; Ruíz-Bustos, 1976; Ruíz-Bustos, 1991; Cuevas et al., 1984 ; Agustí, 1985 ; Mazo et al., 1985 ; Soria, 1986 ; Moyà-Solà et al., 1987; Agustí et al, 1987 ; Martín-Suárez, 1988 ; Alberdi and Bonadonna, 1989 ; Sesé, 1989 , 1991 , 2006 ; Martínez-Navarro, 1991 ; Soria and Ruiz-Bustos, 1991 , 1992 ; Guerra-Merchán, 1992 ; Sesé 1994 ; Sesé and Sevilla, 1996 ; Gibert and Gibert, 2003 ; Ros-Montoya, 2010 ; Maldonado-Garrido et al, 2017 ; Ros-Montoya et al., 2017 , 2018 ; Martínez-Navarro et al., 2018 ). Thirty-three of these sites were selected by their archaeological or paleontological relevance, as well as by their clear geographic, stratigraphic and chronological definition (Table 1 ). Of these sites, 82% (27 out of 33) lie in the Orce sub-basin. Thirteen sites (39%) yielded macrovertebrate remains, with evidence of hominin presence in four of them. These sites were also positioned in a stratigraphic series (Fig. 2 ). In terms of age, 27% of sites are from the Ruscinian, 6% from the Early Villafranchian, 18% from the Middle Villafranchian, 43% from the Late Villafranchian, 3% from the Epivillafranchian, and the remaining 3% from the Galerian. The field study showed the presence of 17 levels of cherts, all located in the Orce sub-basin, mainly to the south of the town of Galera (Fig. 1 C). Some paleontological sites (e.g., FN-1, GA-2 and NU-2) are situated just a few decimeters above these siliceous deposits (Fig. 2 ). Chert beds typically occur in irregular shaped strata, with a lateral extent ranging from 7 m to > 200 m, and a thickness of 8–80 cm (Figs. 3 A, C). The cartographic relationship between the appearance of cherts and the presence of tectonic fractures is remarkable (Fig. 1 C; Figs. 3 G, I, J). Lithologically, these cherts appear as white, dark gray or light brown massive facies. At an outcrop scale, they show abundant diaclases, micropores, changes of coloration, and the presence of conchoidal fractures. In some cases, the cherts appear with decimeter to centimeter-scale gypsum mineralizations within them (Fig. 3 H). Another remarkable aspect is that these cherts appear in different sedimentary contexts, in stratigraphic contact with other six types of facies: limestones, marls, gypsum, (calci)lutites, and sandstones (Fig. 2 ). 3.1 Elemental analysis Quantitative analyses of chemical elements were performed (Tables 3 and 4 ). The results were plotted in the Al-Fe-Mn diagram and the two following ratios were calculated: (Fe + Mn)/Ti and Al/(Al + Fe + Mn) (Fig. 4 and Table 5 ). A common feature of hydrothermal deposits is an Al/(Al + Fe + Mn) ratio of 15 (Boström et al., 1973 ; Murray, 1994 ; Zhang et al., 2023 ). Forty-two out of the fifty-six samples analyzed did not show mineral and/or geochemical indicators that could a priori be linked to hydrothermal contexts, while 14 provided preliminary results that are compatible with a hydrothermal origin according to the Al-Fe-Mn diagram and the (Fe + Mn)/Ti - Al/(Al + Fe + Mn) ratios (Boström et al., 1973 ; Adachi et al., 1986 ; Yamamoto, 1987 ; Murray, 1994 ; Yamamoto et al., 2013 , Fig. 4 and Table 5 ). Eight of these samples (AQ-2, GA-3A, GA-3B, GA-6, GA-7, SX-A, FN1A and CN-1) correspond to cherts, three to travertines (CCH-B, TCV and TE100), two to calcilutites (PL-1 and OR-1) and one to dark lutites (BZ1-A). Except for the dark lutites of the Pliocene site of Baza-1 (Figs. 1 B, 2 ), all the others come from materials in the Orce sub-basin of Late Pliocene to Late Pleistocene age. The (Fe + Mn)/Ti and Al/(Al + Fe + Mn) ratios of these samples are in agreement with conditions of hydrothermalism for the samples of travertines and calcilutites, but not for sample the dark lutites. The last sample provided results of 13.24 and 0.42, respectively. Although these values are very close to the limit of hydrothermal conditions (> 15 and < 0.4, respectively), they are placed outside the area of hydrothermalism. These geochemical ratios could not be applied to chert samples, because neither Al nor Ti were measured in them. Chert samples showed SiO 2 concentrations in excess of 90%, with the exception of two samples, FN1A and GA-6, which exhibited higher concentrations of other compounds (e.g., 14.23% of CaO in FN1A and 5.72% in GA-6). Their variance in composition may be attributed to the presence of calcite in the case of FN1A and gypsum in the case of GA-6, in which a concentration of 6.6% of SO 3 was measured (Table 3 ). Other trace elements are present in proportions < 1% by weight. The geochemical composition of the cherts exhibits an uneven distribution of trace elements (Table 4 ). Al was absent in all cases, and this was also the case for Mn except for samples CN1 and FN1A, in which this element was recorded with abundances of < 0.03%. Fe, Ca, and Cl were consistently detected in all samples. Moreover, Mg, S, K, and Cu were present in every sample except CN1, but the proportions of these elements were lower than in the other samples analyzed in this study. The five additional samples linked a priori to a hydrothermal origin include travertines, dark lutites and calcilutites. The travertine levels (samples CCH-B, TCV and TE100, Fig. 1 C) are located to the west of the town of Orce. They consist of brown stratiform layers (Fig. 3 E) in sedimentary unconformity with the materials deposited below them, which form patchy structures with a lateral development limited to a few meters or, in the case of samples TCV and TE100, to ∼25 meters. Internal annular carbonate growths developed over previous vegetal structures are visible in these travertines (Fig. 3 F). Pisolitic gravels of chaotic aspect and centimeter-scale speleothems are also common. The other travertines present in the Baza Basin show stratiform levels of decimetric thickness in the Venta Micena sector (Fig. 2 ), where they are stratigraphically close to the paleontological sites of this area, and there are also travertine buildings of metric thickness in the vicinity of currently active hydrothermal springs in the Zújar sector (Fig. 1 B). According to the dating of similar travertine deposits in the Alicún de las Torres current hot spring, which is located in the Guadix Basin (Hernández et al., 2000 ; Crespo et al., 2014 ), the age of these travertines is Middle to Late Pleistocene. The dark lutites that show a geochemical profile compatible with a hydrothermal origin consist of a ~ 5 cm thick level that is located at 45 cm from the top of the Baza-1 paleontological site (Figs. 2 , 3 D). From a lithological point of view, these lutites show dark gray tones and contain sub-millimeter-scale parallel laminations, root remains, small fragments of gastropod shells, oxidations, and micas randomly distributed in the sediment. The presence of dark lutites in the Baza Basin is commonly recorded in thin layers present in most of the stratigraphic series of Plio-Pleistocene age (Fig. 2 ). Mesoscopic scale sedimentary structures of traction origin and linked to detrital deposits (e.g., sandy microlevels, coarsening/finning upward sequences, imbrications, or scour-marks) are absent. Analyses of sample BZ1-A (Tables 3 – 4 ) showed a main chemical composition of Si > Fe > Al > Ca > K > Mg. Compared to other dark lutites studied, BZ1-A stands out for its notably higher concentration of Fe 2 O 3 , which reaches 11.19%. This iron content exceeds those of other samples of dark lutites. Additionally, this sample exhibits elevated levels of K, Mn, S, Cl, Ba, Cu, Rb, Pb, Sn, and Cs compared to the average contents of the samples of dark lutites. Calcilutites are present in all stratigraphic profiles except Zújar (Fig. 2 ) and use to appear as light-colored levels of decimetric to metric thickness. From a lithological point of view, they typically exhibit massive facies or scarce internal arrangement, although sometimes there are internal shallowing-upward sequences with edaphogenic-type facies of reddish tones situated at their top. The PL-1 calcilutite sample shows a main geochemical composition of Ca > Si > Fe > Al > Mg (Tables 3 – 4 ). The rest of the elements detected appeared with concentrations < 0.6%. In comparison with other calcilutites of the basin, this sample has the highest CaO values and the lowest of SiO 2 , Al 2 O 3 , K 2 O, TiO 2 , Na 2 O, and SO 3 . Finally, the OR-1 calcilutite sample shows a main geochemical composition of Si > Ca > Mn > Al > Fe > Mg. The rest of the elements detected appear with concentrations < 0.7%. This sample exhibits the highest MnO value (~ 10 times more) compared to the average of the calcilutite samples. Additionally, geochemical analyses revealed the presence of some trace elements in the samples (Table 4 ). Of these, Ba stands out, being exclusively detected in four samples a priori linked to hydrothermal environments: BZ1-A (dark lutites), OR-1 (calcilutites), TCV (travertines), and TE100 (travertines). 3.2 Mineralogical analysis The XRD data of cherts show that opal is the principal phase (Table 6 , Fig. 5 ). The d (101) (> 4.07 Å) and the full width at half-high maximum (FWHM) reflection peak intensity of opal (101) indicate that CT is the polymorph of the opal (Elzea et al., 1994 ). Most of the samples taken in Galera showed the presence of gypsum in the XRD data, as evidenced by geochemistry (high CaO and SO 3 contents, Table 3 ) and field observations (Fig. 3 H). The presence of silicates and carbonates in some samples (e.g., CN-1, GA-3B and SX-A) may come from the remains of the sedimentary materials above and/or below the levels where these samples were obtained, due to the irregular morphology of the chert strata. Travertine samples mainly consist of calcite (Table 6 ) and, to a lesser extent, pyrolusite. In addition, other mineral species such as gypsum and halite were found. The XRD data from these samples agree with the expectations from geochemical (Tables 3 – 4 ) and lithological data. The analyses carried out on the calcilutites of samples PL-1 and OR-1 (Table 6 ) showed a mineral composition of calcite, quartz and phyllosilicates (muscovite and paragonite). In addition, dolomite and plagioclase were both present as trace amounts. Finally, the geochemistry of the sample of dark lutites with hydrothermal markers (BZ1-A) consisted mainly of phyllosilicates (> 50%: muscovite, paragonite, kaolinite, and chlorite) and quartz (Table 6 ). Accessory minerals included feldspars, carbonates, lepidocrocite, halite, and gypsum. The high FeO 3 , CaO, SO 3 , and Cl (Tables 3 and 4 ) contents justified the presence of lepidocrocite, gypsum and halite. 4 Discussion On a cartographic scale, 14 samples linked to hydrothermal environments were found in the vicinity of traces of the main tectonic fractures present in the basin (Sanz de Galdeano et al., 2007, 2012; García-Tortosa et al., 2011 ; Figs. 1 B–C): samples AQ-2, GA-3A, GA-3B, GA-6, GA-7, and PL-1 are close to the NE-SW Galera Fault; sample CN1 is close to the NW-SE Alfahuara-Botardo Fault; sample BZ-1A is close to the NNW-SSE Baza Fault; and samples SX-A, FN1A, OR-1, CCH-B, TCV, and TE100 are all in the vicinity of WNW-ESE oriented fractures. This suggests the connection between materials with evidence of hydrothermal activity and their position close to tectonic fractures, as evidenced by the fact that the current thermal springs in the basin are found on or in the vicinity of traces of tectonic fractures (Fig. 1 B). At an outcrop scale, the relationship between chert deposits and tectonic fractures is evident (Figs. 3 G, I, J). 4.1 Origin of cherts and other facies potentially generated in hydrothermal contexts Chert layers appear in the NE sector of the Baza Basin (Orce sub-basin) and are cartographically associated with fracture traces. In fact, a chert dyke (Fig. 3 I) was recognized to the SE of the town of Galera, coincident with the trace of a NE-SW fault that shows synsedimentary features (Sánchez-Roa et al., 2016 ). In the case of Fuente Nueva-1 and Galera, the chert deposits are found near levels rich in paleontological remains (Fig. 2 ). Hydrothermal deposits often show high concentrations of iron and manganese, while aluminum and titanium are reliable indicators of sedimentary input (Boström et al., 1973 ; Adachi et al., 1986 ; Murray, 1994 ; Zhang et al., 2023 ). This is why many research studies use the ratios (Fe + Mn)/Ti and Al/(Al + Fe + Mn) to identify hydrothermal influence (Table 5 ). The application of the Al-Fe-Mn diagram (Adachi et al., 1986 ; Yamamoto, 1987 ; Yamamoto et al., 2013 ) to the chert samples (Fig. 4 ) showed a geochemical composition in agreement with a hydrothermal origin. Although Al-Fe-Mn diagrams have been applied to deposits of marine origin (Adachi et al., 1986 ; Yamamoto, 1987 ; Boncheva et al., 2023 ; El-Moghazy et al., 2023 ; Jurkowska and Swierczewska-Gladysz, 2024; among others), they have also been applied to continental environments (Madondo et al., 2021 ; Zhou et al., 2022 ). The results obtained in the analysis of the chert samples from the Baza Basin are compatible with a hydrothermal origin for them if we consider other evidence (see below) and also according to the Al-Fe-Mn contents and main trace elements present in the siliceous rocks of the North Pacific (Adachi et al., 1986 ). On the opposite, the (Fe + Mn)/Ti and Al/(Al + Fe + Mn) ratios have been used in sediments to determine their hydrothermal origin (Boström et al., 1973 ; Murray, 1994 ; Zhang et al., 2023 ). Therefore, the data obtained from the chert samples are expected to be more reliable than those derived from samples without chert contents. In any case, the use of both methods is complementary and the comparison of their results in these continental environments allows their evaluation as proxies in this sedimentary context. The main mineral composition of opal-CT in the chert samples indicates a silica mineral phase linked to hydrothermal conditions (Kano and Taguchi, 1982 ; Flörke et al., 1990 ; Lynne et al., 2005 ; Fröhlich, 2020 ). Silica-rich deposits linked to > 250ºC hydrothermal fluids have been associated with tectonic fractures in Permian cherts based on their geochemical analyses and Al-Fe-Mn diagrams (for review and references, see Zhou et al., 2022 ). According to the Na/K calculation method (Fournier and Truesdell, 1973 ; Fournier, 1977 ; Giggenbach et al., 1988 ), the estimates obtained with geothermometers for the high-temperature hot springs that are currently active in the Guadix-Baza Depression are 88–89ºC. A similar situation occurs in Alhama de Granada, SE Spain (López-Chicano et al., 2001 ). Moreover, the hot springs of the Guadix-Baza Depression (Diputación de Granada-ITGE, 1990 ; Table 2 ) show a direct relationship between their output water temperature and silica contents (García-Aguilar et al., 2014 ), which can reach up to 54 mg/l at 44º C in the Graena spring. According to the standard values of the upper crustal geothermal gradient, tectonic and geophysical gravimetry data, Sanz de Galdeano et al. (2007) suggested a temperature range in the deep traces of the main faults of the Baza Basin of 150–300ºC, which agrees with a hydrothermal scenario. These temperatures are compatible with those inferred for the formation of Permian chert nodules in the Junggar Basin (Zhou et al., 2022 ). The trace-element contents of the cherts of the Baza Basin are very variable depending on the sample considered, because each of them shows a particular geochemical profile. The case of sample FN1A, which is located a few decimeters below the Fuente Nueva-1 site, stands out for its variety of trace elements, many of them not recorded in other deposits (e.g., Sr, As, Mo, and W; Table 4 ). In this sense, the presence of As has been associated with hydrothermal fluids near tectonic fractures west of the Orce sub-basin (Sánchez-Roa et al., 2016 ) and the presence of Sr has also been linked to hydrothermal contexts in the Plio-Pleistocene lacustrine environments of the Guadix-Baza Depression (García-Aguilar et al., 2014 ). On the other hand, the variety of geochemical profiles found in the chert samples is compatible with the variety of hydrochemical profiles (both qualitatively and quantitatively) and output temperatures present in the hot springs that are currently active in the basin (Table 2 ). These hydrochemical profiles must be linked to the geometry of the tectonic fractures associated with each hot spring and to the different lithological substrates that the hydrothermal fluids passed through on their way to the surface. This cause-effect relationship could be extrapolated to the ancient Plio-Pleistocene hydrothermal upwellings in the Baza Basin, whose salt contents, output temperatures and water flow would have varied depending on the tectonic, lithological, and structural features present in each case. The hydrochemical composition of the hot springs that are currently active in the basin with output temperatures of > 35ºC (Cruz-Sanjulián et al., 1972 ; Cruz-Sanjulián and García-Rosell, 1972 , 1975 ; Diputación de Granada-ITGE, 1990 ; Table 2 ) shows silica contents of > 40 mg/l, high sulfate values and the presence of chlorides. Similar conditions are widely documented in hydrothermally active regions, such as the Dead Sea-Jordan River Basin in Israel (Levitte and Eckstein, 1978 ), Yellowstone in USA (Nordstrom et al., 2005 ), Mount Tafta in Iran (Shakeri et al., 2008 ) or Denizli in Turkey (Özler, 2000 ), among others. On the other hand, White et al. ( 1956 ) and Fournier and Rowe ( 1966 ) established the hydrochemical conditions linked to the solubility and subsequent precipitation of silica in thermal waters. They defined a solubility of 315 mg/l for silica at 90ºC and 110 mg/l at 25º C, demonstrating a correspondence between the output temperature of the hot springs and their silica contents. The rapid rise of hot waters to the surface results in silica oversaturation due to cooling and degasification. Dissolution by high-temperature waters of deep rocks and their subsequent ascent through tectonic fractures is common. In the case of cherts, however, these hot springs should have shown much higher output temperatures (> 35ºC) than in the atmosphere. Considering the climatic conditions inferred for the Early Pleistocene in the Baza Basin, these thermal pools would have originated small areas with a stable and warm microclimate throughout the year, suitable for the ecological development of a diverse mammalian community, especially during the cold periods of the Pleistocene (Lisiecki and Raymo, 2007 ; Paillard, 2017 ). Under these conditions, cherts originated from the dissolution at depth conditions of rocks with elevated siliceous contents (e.g., quartzites and mica-schists of the Internal Zones of the Betic mountain range; Baena Pérez et al., 1979 ; Martín García et al., 1980 ; Villalobos et al., 2006 ) in high-temperature fluids. According to the most plausible calculated Na/K geothermometers (88–89ºC) and considering the standard geothermal gradient in the upper crust (25–30ºC/km, Harrison et al., 1990 ; Gupta and Roy, 2007 ), the estimated depth in the Baza Basin for the dissolution of silica agrees with the basement depth calculated from seismic profiles, 2150 m (Alfaro et al., 2008 ). This depth is similar to the estimation for present-day hot springs in Algeria (Belhai et al., 2017 ) and indicates the presence of a deep fracture network affecting the basin substrate in the Guadix-Baza Depression, where the hot springs would dissolve components of the rocks bound to them. These depth values linked to the origin of thermal upwellings in the basin are lower than the depth range of the tectonic fractures in the basin (5–10 km), which has been estimated using geological and geophysical criteria (Sanz de Galdeano et al. 2012, 2020). Travertine levels of decimetric thickness and associated with a hydrothermal context have been detected in the southern bank of the Orce River, west to the town of Orce. The application of the Al-Fe-Mn diagram and the (Fe + Mn)/Ti and Al/(Al + Fe + Mn) ratios to these samples showed a composition compatible with a hydrothermal origin (Fig. 4 , Table 5 ). Furthermore, the Al-Fe-Mn diagram suggests that sample BZ1-A of dark lutites may have had some hydrothermal influence. Moreover, the mineralogy of this sample (Table 6 ) indicates a primarily detrital deposit (quartz, paragonite and chlorite) with halite precipitation due to evaporation processes. It is noteworthy that the presence of Sr and Mo was detected in this sample (Table 4 ), as these elements have been found under a hydrothermal context in the Galera deposits (Sánchez-Roa et al., 2016 ). Despite having very high iron contents, the hydrothermal ratios (Fe + Mn)/Ti and Al/(Al + Fe + Mn)) suggest that this level has not a hydrothermal origin. Given the discrepancy between the Al-Fe-Mn diagram and these ratios, it is important to note that the diagram was designed for analyzing siliceous materials (Adachi et al., 1986 ) while the ratios are typically used for sediments (You et al., 2020 ; Zhang et al., 2023 ). As a result, it is plausible that the ratios provide a more accurate indication of hydrothermalism in these dark lutites. However, although the values in these ratios are close to those expected in hydrothermal environments, they are not within the hydrothermal zone. This casts doubts on whether BZ1-A was influenced by hydrothermalism. In any case, it is remarkable that BZ1-A is the only sample among the 17 samples of dark lutites analyzed that fits the diagram of hydrothermalism. The calcilutite samples that provided geochemical indicators of hydrothermal activity correspond to two metric scale levels located near the localities of Huéscar (sample PL-1) and Orce (sample OR-1, Fig. 2 ). In both cases, they are placed in the vicinity of paleontological sites with macrofauna. The application of the (Fe + Mn)/Ti and Al/(Al + Fe + Mn) ratios to these samples (Table 5 ) shows values that are compatible with a hydrothermal origin for them. In the case of sample PL-1, the Al-Fe-Mn diagram (Fig. 4 ) shows further evidence of its hydrothermal origin. From the mineralogical point of view, the composition of these samples shows quartz, calcite, dolomite, muscovite, paragonite, and plagioclase (Table 6 ). This association of minerals can be considered detrital-inherited by sedimentary processes, as evidenced by the lithological characteristics observed in the outcrops where the samples were taken. In addition to the previous arguments, the presence of significant amounts of Ba in samples BZ1-A (dark lutites), OR-1 (calcilutites), TCV (travertines), and TE100 (travertines) adds further evidence on the hydrothermal context associated with their deposition (Jankovic, 1982; Canet et al., 2005 ). The hydrothermal markers in samples Pl-1, OR-1 and BZ1-A, whose lithologies contain inherited detrital elements, could be interpreted as due to minerals originating from source areas with metamorphic substrates. However, these hydrothermal markers do not appear in other deposits that are lithologically and sedimentologically similar to these three samples, which indicates that the high Fe and Mn contents should have a hydrothermal origin. Concerning the sedimentary context of the cherts and other deposits linked to hydrothermal conditions and their association with archaeological and paleontological sites, it is worth noting that these levels frequently appear on the top and/or bottom of the sites as well as in lateral connection with facies linked to a diversity of sedimentary scenarios. 4.2 Paleogeographical reconstruction of hydrothermal activity in the basin The spatio-temporal correlation between the coordinates of the 14 samples with evidence of past hydrothermal activity and the position of the hot springs that are currently active in the Baza Basin allows us to propose a general model of paleogeographic evolution of the basin in relation to its hydrothermal activity (Fig. 6 ). This model makes it possible to specify the cause-effect relationships between the tectonic dynamism of the basin and the occurrence of hydrothermal phenomena linked to tectonic fractures (Figs. 1 B–C) at specific places and times that generated in each case different geochemical and mineral markers in the sedimentary environments. These environments can be very diverse, since they do not show a direct genetic relationship with the position of the hydrothermal upwellings. In this way, the cherts, travertines, calcilutites, and dark lutites (i.e., the facies in which hydrothermal markers have been detected) correspond to different sedimentary environments, which include lacustrine zones with some depth (calcilutites), marshy environments (dark lutites), occasional upwellings in favor of tectonic fractures (cherts) and waterfalls or pools of thermal waters (travertines), according with the most plausible sedimentary models (Reading, 1978 ; Prothero and Schwab, 2004 ). Among these sedimentary environments, those linked to the cherts could be considered the only ones with a direct causal relationship between these facies and the presence of hydrothermal activity. As shown in Fig. 6 , hydrothermal phenomena appear discontinuously in the Baza Basin from the Late Pliocene (Ruscinian) to the present. In this sense, there seems to be a gap between the Upper Ruscinian and the Middle Villafranchian, together with a gap in the lower Late Villafranchian. However, the presence of hydrothermal activity during these periods cannot be ruled out. Its demonstration would depend on a wide sampling of the different sedimentary facies linked to these periods, preferably located near tectonic fractures. Regarding the geographical distribution, it is evident that hydrothermal activity concentrates in the Galera-Orce-Cañada del Negro sector (Orce sub-basin). This points to an intense tectonic activity in this sub-basin during the Plio-Pleistocene, particularly in four intervals: Late Ruscinian, Middle Villafranchian, upper Late Villafranchian and Galerian to present, with the presence of hydrothermal phenomena that induced mineral and/or geochemical markers in the sedimentary environments associated to these places and times. Likewise, the Baza area seems to show a similar pattern to the Orce sub-basin for the Ruscinian, associated with an important tectonic fracture, the Baza Fault. It is also evident that each node of hydrothermal activity seems to demonstrate a particular physico-chemical characteristic, as shown by the compositional variety of the deposits linked to this activity. In this regard, factors such as the hydrochemical composition and physical parameters of the hydrothermal fluids, the sedimentary environments where they acted and (micro)biological factors would give rise to a wide range of situations. This pattern could have played an important role in the development of local ecosystems in the vicinity of the nodes of hydrothermal activity. Finally, it is evident how these hydrothermal phenomena coincide quite closely in their spatio-temporal coordinates with the position of important paleontological sites, as happens in the case of BZ-1, FN1, HU-1, and BO-7 (Fig. 2 ). If we further extend these spatio-temporal coordinates to a wider range, we see other important sites (e.g., BL, FN-3 and VM) that appear in a geographic and/or chronostratigraphic position close to the nodes of hydrothermal activity. This allows us to establish a possible paleoecological relationship between the presence of hydrothermal phenomena and the occurrence of paleontological sites in the Baza Basin. 4.3 Paleoecological implications of hydrothermal activity in the basin A contribution of thermal waters would play a role in maintaining during the dry season a minimum level in the water table of the lacustrine environments of the Baza Basin. This would in turn allow the development of a luxurious vegetational cover in the surroundings of the hot springs, which would sustain a large vertebrate community. The hot springs of the Baza Basin led to the appearance of thermal pools of variable extent and length, with waters that differed from the main lacustrine systems in their composition and physical properties. These hot springs would support in part the ecological pyramid of the Baza Basin in a specific network of spatio-temporal nodes, concentrating the plants and animals in their surroundings, which would in turn result in a concentration of ungulate carcasses in the vicinity of the hot springs (Fig. 7 ). This is reflected in some of the most important deposits of the basin, such as the Pliocene sites of Baza-1 and Fuente Nueva-1, which are both linked to a level of dark lutites and to a level of cherts, respectively. From a sedimentary point of view, this provides evidence on the role of hydrothermal events as hot spots for the concentration of terrestrial fauna. The relationship between some ecological factors linked to contexts with thermal waters is shown in Fig. 8 A, which depicts a direct, statistically significant relationship between prey biomass (kg/km 2 ) and annual rainfall (mm) values in a set of African Natural Parks and Game Reserves [data compiled from Bell ( 1981 ) and Hatton et al. ( 2015 ); see also Coe et al. ( 1976 ) for a similar relationship]. However, there is considerable scatter around the least-squares regression line, which primarily depends on soil type and, secondarily, on the presence of thermal springs (Fig. 8 B), which increase the net primary productivity thanks to their saline contents (García-Aguilar et al., 2014 , 2015 ; Palmqvist et al., 2022a ). Figure 8 also includes the mean estimates of carrying capacity for the Orce sites (3827.8 kg/km 2 for VM and 3307.6 kg/km 2 for BL/FN-3), obtained using an approach based on the Weibull model (see Rodríguez-Gómez et al., 2022 , 2024 ), and the estimates of annual rainfall, based on the composition of the herpetological assemblages in the case of BL/FN-3 (700 mm for BL and 738 mm for FN-3; Blain et al., 2016 ) and on the abundance of nitrogen isotopes in the bone collagen retrieved from the fossils of large mammals from VM (800 mm; Palmqvist et al., 2022c , 2023 ). As deduced for the Baza Basin, the presence of thermal waters provided a variety of salts and trace elements. These mineral nutrients allowed the development of a diverse plant community and the translation of this biodiversity to the local trophic pyramid, thus configuring a sort of ecological "Eden" on a limited spatio-temporal scale. Given that some species of mammals, for example the hippopotamuses, used these thermal pools and watered them, their salinity (at least where the fossil remains are found) must have been of < 5 g/l (Palmqvist et al., 2022a ). In this environmental context, the ancient hominin settlers of the Baza Basin sought such places as preferential sources of water and a variety of trophic resources. Figure 8 B shows that although the productivity of the African National Parks and Game Reserves mainly depends on annual precipitation, as noted above, it also relates to soil type and to the presence of thermal waters, which contribute to increase the productivity of the terrestrial ecosystems and their herbivore biomass. Specifically, the areas with volcanic and Rift sediments have more positive residuals in the regression equation of Fig. 8 A than those with other types of sediments (i.e., basement, floodplain, granite, Kalahari sands, and marine sediments; see also Bell, 1981 ). Similarly, the areas with active hot springs show greater values of prey biomass than those in which thermal activity is absent (Fig. 8 B). 4.4 Correlation of the proposed model for the Baza Basin with other World sites A large-scale geographical position of 103 African, West Asian and European sites, which range in age from 7.5 to 0.35 Ma, shows a close relationship with those areas that show high tectonic and geothermal activity (Fig. 9 ). Most of these sites appear in areas that evidence intense geological activity related to seismotectonic, volcanic and hydrothermal phenomena during the Plio-Pleistocene, and even up to present days. These sites usually show tuffaceous levels, volcanic ashes, seismites, or lacustrine deposits potentially fed by thermal waters. This is also the case of the archaeological deposits of Java (Indonesia), where volcanism and seismicity are common and particularly intense (Delinom, 2009 ). These sites range in age from the appearance of the first hominins in Africa to the arrival of Homo sapiens in Morocco, as detected in the Jebel Irhoud site (Hublin et al., 2017 ). Figure 9 shows that 88% of the sites are found in areas where volcanic, seismic, tectonic, and hydrothermal phenomena have been common since Late Miocene times (and in many cases up to present days). Of all these sites, 71% appear in sedimentary facies of lacustrine or fluvio-lacustrine origin, as happens throughout the East African Rift, while the other sites are found in karst cavities (21%) and in sedimentary formations of fluvial or marine-littoral origin (8%). Most paleoanthropological sites placed in geologically stable areas are found in karst cavities and fluvial paleo-terraces. If we consider the sedimentary origin and the tectonic environment of these sites, 67% of them are found in lacustrine or fluvio-lacustrine facies located in tectonically active areas with hot springs. The network of spatio-temporal nodes of hydrothermal phenomena in the Baza Basin seems to be related with the presence of hominin settlements, as evidenced by four archaeological localities: the late Early Pleistocene sites of Barranco León and Fuente Nueva-3, the Early-Middle Pleistocene site of Huéscar-1, and the early Middle Pleistocene site of Cúllar-Baza-1. This possible association has led us to the proposal of a new idea on the distribution of paleoanthropological sites on a global scale during the Plio-Pleistocene. Many African and Eurasian sites appear in areas that evidence intense geological activity related to seismotectonic, volcanic and hydrothermal phenomena during the Plio-Pleistocene. In most cases, these areas remain tectonically active today (Fig. 9 ). Such scenarios can be compared with those that characterized the Plio-Pleistocene environments of the Baza Basin. Throughout the East African Rift Valley, from Mozambique-Malawi to Ethiopia and Eritrea, following the Red Sea and the Levantine Corridor to Anatolia and Georgia, the main archaeological deposits are found in volcanic contexts and scenarios characterized by intense tectonic activity, which resulted in geothermal anomalies during the Neogene-Quaternary (Morley et al., 1992 ; Schlüter, 1997 ; Adamia, 2013 ; Glerum et al., 2020 ). More specifically, a relationship between hydrothermal activity and paleoanthropological deposits of 1.7 Ma has been evidenced by microbial markers in Olduvai Gorge by Sistiaga et al. ( 2020 ), who used a similar model to the one proposed here for the Baza Basin. Such correspondence also occurs in several European sites linked to recent hydrothermal and seismo-tectonic activity, for example the Acheulean site of Rodafnidia, Lesvos Island (Galanidou et al., 2013 , 2016 ). This shows the common relationship between areas with intense geothermal activity and the presence of archaeological sites with hominin remains and fossils of large mammals. Tectonic activity is a source of geomorphological variability (i.e., landscape roughness) and ecological diversity (Coppens, 1994 ; Foley, 1987 ; Gamble, 1993 ) and has been linked to the first hominins that evolved in the African Rift, because it allowed them to obtain stable resources: the tectonic and volcanic movements that generated the East African Rift System resulted in a broken, hilly rough country of complex topography where the early hominins could hunt and hide (Bailey et al., 2000 , 2011 ). Moreover, environmental roughness probably characterized the whole route of early hominin dispersal (King and Bailey, 2006 ). We suggest here that tectonic and hydrothermal activity are significantly linked to the evolution of the human lineage due to the consumption strategies of hominins and their risk of exposure to confrontation with carnivores. In fact, carnivores could have predated the own hominins (Treves and Palmqvist, 2007 ) and were also competitors in the access to meat, although they were also providers of carrion (Espigares et al., 2013 ; Palmqvist et al., 2022a , 2023 ). According to the evidence unearthed from the fossil sites of Orce, the hominins developed a scavenging behavior as a successful strategy to obtain animal resources in the Baza Basin (Espigares et al., 2019 ; Palmqvist et al., 2023 ). Analyses on resource partitioning among the members of the carnivore guild suggest that scavenging was the optimal strategy for these hominin populations (Rodríguez-Gómez et al., 2016 ). Anatomical evidence from late Early Pleistocene sites younger than those of Orce, such as level TD6 from Gran Dolina in Sierra de Atapuerca, suggests that these humans had lesser throwing skills than modern humans, which would hinder a hunting activity based on throwing cobbles against prey, although it would not have limited other hunting strategies (Bermúdez de Castro et al., 2020; however, for an opposite interpretation see García-Martínez et al., 2021 ). A heterogeneous environment would be useful for exploiting different ecological niches in the case of an opportunistic species like Homo sp., relatively weak for a direct confrontation with the large carnivores. Moreover, these mosaic habitats would also provide protection from cursor predators in the open environments and from ambushing carnivores in the closed ones, as the hominins could inhabit the boundary zones between different environments, both for protecting themselves from predators and for exploiting the resources of these environments (see Treves and Palmqvist, 2007 ). In ecology, the transition zones between different ecosystems are called ecotones. We propose that rather than a generalist species, the early members of Homo would be specialized in exploiting the ecotones, thus showing adaptations to consume resources from different environments. According to an analysis of avifauna in Lower, Middle and Upper Paleolithic sites (Finlayson et al., 2011 ), there is a significant association between hominin presence and mosaic habitats (i.e., habitats with many ecotones). On the other hand, hominin absence of Sierra de Atapuerca during the early Middle Pleistocene has been linked to a greater environmental homogeneity than during previous periods (Rodríguez et al., 2011 ; Rodríguez-Gómez et al., 2014 ). Today, the areas with highest ecosystem diversity are two regions with active tectonics, the African Rift and the Himalayan range, in accordance with the ecoregions defined by Olson et al. ( 2001 ). The richness of ecoregions per unit area observed in the African Rift is probably lower nowadays than during the Plio-Pleistocene, because tectonic activity was higher at that time (Bailey et al., 2000 ; King and Bailey, 2006 ). This diversity of ecoregions seems to relate to the diversity in the values of abiotic and biotic variables that can be observed in the region. For these reasons, the African Rift would provide a high diversity of habitats in which a generalist or ecotone-specialist species, such as early Homo , could have efficiently exploited these environments for resources and shelter. This model can be proposed for the ancient hominin populations found in the archaeological sites of the Baza Basin (for a review on the paleodemography of these populations, see Palmqvist et al., 2022a ). From a paleoecological point of view, the Ngorongoro Crater in the African Rift Valley is a model that can be correlated with the sites of the Baza Basin and other deposits found in geologically active areas. In Ngorongoro, the hydrothermal influence of the crater lakes has a seasonal pattern. As a result, the populations of large mammals are dispersed in the wet season, while during the dry season they concentrate near the pastures that surround the hot springs. These waters lose their ecological influence at ~ 700 m from their point of emergence, changing to alkaline (pH = ~ 8) waters with high saline contents (DeoCampo, 2002 ). The list of species that live in the crater within a radius of ~ 100 m around the thermal springs includes buffaloes, gazelles, hippopotamuses, rhinoceroses, zebras, elephants, antelopes, wild boars, hyenas, foxes, cheetahs, jackals, and lions. This fauna is strikingly similar to the one that inhabited the Baza Basin during the Plio-Pleistocene in terms of their distribution among modes of locomotion, feeding types and body size classes (Mendoza et al., 2005 ). 5 Conclusions Hot springs have been important for faunal and hominin survival during the Plio-Pleistocene, favoring the dispersal of mammalian species as documented in the Baza Basin, which preserves one of the continental sequences with greater stratigraphic completeness of Europe, including the earliest archaeological evidence of hominin presence in the northern Mediterranean latitudes during the late Early Pleistocene. In this basin, hot springs form a network of spatio-temporal nodes, which are particularly concentrated in the Orce sub-basin, showing permanent siliceous waters with temperatures > 35º C. These hot springs played an important ecological role during the summer, when seasonal rain-fed springs became dry, and also helped to maintain locally warm temperatures in winter, thus aiding the survival of water dependent vertebrates adapted to warm and moist climates, including the members of early Homo. Nearly one hundred Plio-Pleistocene archaeological and paleontological localities are recorded in the perimeter of the Baza Basin. Most of these sites were generated in connection with tectono-sedimentary processes, as evidenced by the presence of paleo-seismites, ancient hot springs and faulting. The lithological, mineralogical and geochemical composition of the chert levels present in the Orce sub-basin suggests a high-temperature hydrothermal origin for these deposits. Other types of sedimentary facies present in the basin, such as travertines, dark lutites or calcilutites, were influenced by this network of spatio-temporal nodes of hot springs, showing diverse geochemical and mineralogical indicators associated with this hydrothermal origin. The mineral nutrients provided by the hot springs prompted the development of a luxurious vegetation, which in turn allowed the development of a diverse community of large mammals, including the first hominins that dispersed in Western Europe. Declarations Compliance with Ethical Standards Disclosure of potential conflicts of interest: On behalf of all authors, the corresponding author states that there is no conflict of interest. Research involving Human Participants and/or Animals: This research involved no human participants and/or animals. 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Sedimentary basin destruction inferred from the evolution of drainage systems in the Betic Cordillera, southern Spain. Journal Of The Geological Society Of London , 149 , 1021–1029. https://doi.org/10.1144/gsjgs.149.6.1021 . Warr, L. N. (2021). IMA–CNMNC approved mineral symbols. Mineralog Mag , 85 (3), 291–320. https://doi.org/10.1180/mgm.2021.43 . White, D. E., Brannock, W. W., & Murata, K. J. (1956). Silica in hot-spring waters. Geochim Cosmochim Act , 10 , 27–59. https://doi.org/10.1016/0016-7037(56)90010-2 . White, T. D., Asfaw, B., De Gusta, D., Gilbert, H., Richards, G. D., Suwa, G., et al. (2003). Pleistocene homo sapiens from middle awash, Ethiopia. Nature , 423 , 742–747. https://doi.org/10.1038/nature01669 . White, T. D., Hart, W. K., Walter, R. C., Wolde, G., de Heinzelin, J., Clark, J. D., et al. (1993). New discoveries of Australopithecus at Maka in Ethiopia. Nature , 366 , 261–265. https://doi.org/10.1038/366261a0 . White, T. D., Suwa, G., & Asfaw, B. (1994). Australopithecus ramidus , a new species of early hominid from Aramis, Ethiopia. Nature , 371 , 306–312. https://doi.org/10.1038/371306a0 . Williams, M. A. J., Williams, F. M., Gasse, F., Curtis, G. H., & Adamson, D. A. (1979). Plio–Pleistocene environments at Gadeb prehistoric site, Ethiopia. Nature , 282 , 29–33. https://doi.org/10.1038/282029a0 . Yamamoto, K. (1987). Geochemical characteristics and depositional environments of cherts and associated rocks in the Franciscan and Shimanto Terranes. Sediment Geol , 52 (1–2), 65–108. https://doi.org/10.1016/0037-0738(87)90017-0 . Yamamoto, M., Nakamura, R., Oguri, K., Kawagucci, S., Suzuki, K., Hashimoto, K., et al. (2013). Generation of Electricity and Illumination by an Environmental Fuel Cell in Deep-Sea Hydrothermal Vents. Angewandte Chemie Int Ed , 52 (41), 10758–10761. https://doi.org/10.1002/anie.201302704 . Yeshurun, R., Zaidner, Y., Eisenmann, V., Martínez-Navarro, B., & Bar-Oz, G. (2011). Lower Paleolithic hominin ecology at the fringe of the desert: faunal remains from Bizat Ruhama and Nahal Hesi, Northern Negev. Israel J Hum Evol , 60 , 492–507. https://doi.org/10.1016/j.jhevol.2010.01.008 . You, J., Liu, Y., Zhou, D., Zheng, Q., Vasichenko, K., & Chen, Z. (2020). Activity of hydrothermal fluid at the bottom of a lake and its influence on the development of high-quality source rocks: Triassic Yanchang Formation, southern Ordos Basin, China. Australian Journal Of Earth Sciences , 67 (1), 115–128. https://doi.org/10.1080/08120099.2019.1612783 . Yravedra, J., Solano, J. A., Courtenay, L. A., Saarinen, J., Linares-Matás, G., Luzón, C., et al. (2021). Use of meat resources in the Early Pleistocene assemblages from Fuente Nueva 3 (Orce, Granada, Spain). Archaeol Anthropol Sci , 13 , 213. https://doi.org/10.1007/s12520-021-01461-7 . Yravedra, J., Solano, J. A., Herranz-Rodrigo, D., Linares-Matás, G. J., Saarinen, J., Rodríguez-Alba, J. J., et al. (2022). Unravelling hominin activities in the zooarchaeological assemblage of Barranco León (Orce, Granada, Spain). J Paleolit Archaeol , 5 , 6. https://doi.org/10.1007/s41982-022-00111-1 . Zhang, K., Liu, R., Bai, E., Zhao, Z., Peyrotty, G., Fathy, D., et al. (2023). Biome responses to a hydroclimatic crisis in an Early Cretaceous (Barremian–Aptian) subtropical inland lake ecosystem, Northwest China. Palaeogeogr Palaeoclimatol Palaeoecol , 622 , 111596. https://doi.org/10.1016/j.palaeo.2023.111596 . Zhou, J., Yang, H., Liu, H., & Jiao, Y. (2022). The Depositional Mechanism of Hydrothermal Chert Nodules in a Lacustrine Environment: A Case Study in the Middle Permian Lucaogou Formation, Junggar Basin, Northwest China. Minerals , 12 (10), 1333. https://doi.org/10.3390/min12101333 . Tables Tables 1-6 is available in the Supplementary Files section. Supplementary Files Table1.docx Table2.docx Table3.docx Table4.docx Table5.docx Table6.docx Cite Share Download PDF Status: Under Review Version 1 posted Editorial decision: Minor revisions 28 Apr, 2024 Reviewers agreed at journal 01 Apr, 2024 Reviewers invited by journal 01 Apr, 2024 Editor invited by journal 29 Mar, 2024 Editor assigned by journal 25 Mar, 2024 First submitted to journal 23 Mar, 2024 You are reading this latest preprint version Research Square lets you share your work early, gain feedback from the community, and start making changes to your manuscript prior to peer review in a journal. As a division of Research Square Company, we’re committed to making research communication faster, fairer, and more useful. We do this by developing innovative software and high quality services for the global research community. Our growing team is made up of researchers and industry professionals working together to solve the most critical problems facing scientific publishing. Also discoverable on Platform About Our Team In Review Editorial Policies Advisory Board Help Center Resources Author Services Accessibility API Access RSS feed Manage Cookie Preferences © Research Square 2026 | ISSN 2693-5015 (online) Privacy Policy Terms of Service Do Not Sell My Personal Information {"props":{"pageProps":{"initialData":{"identity":"rs-4151726","acceptedTermsAndConditions":true,"allowDirectSubmit":false,"archivedVersions":[],"articleType":"Research Article","associatedPublications":[],"authors":[{"id":286229074,"identity":"657903d5-079b-4768-ae4c-139b6c1999a4","order_by":0,"name":"José Manuel García-Aguilar","email":"","orcid":"","institution":"University of Malaga: Universidad de Malaga","correspondingAuthor":false,"prefix":"","firstName":"José","middleName":"Manuel","lastName":"García-Aguilar","suffix":""},{"id":286229075,"identity":"8bbc576e-ee29-4417-9bd6-34fd30ca4fea","order_by":1,"name":"Isidoro Campaña","email":"","orcid":"","institution":"University of Malaga: Universidad de Malaga","correspondingAuthor":false,"prefix":"","firstName":"Isidoro","middleName":"","lastName":"Campaña","suffix":""},{"id":286229076,"identity":"7c1cfbf9-0ead-4d66-a2d4-df80e78651cf","order_by":2,"name":"Bienvenido Martínez-Navarro","email":"","orcid":"","institution":"IPHES: Instititut Catala de Paleoecologia Humana i Evolucio Social","correspondingAuthor":false,"prefix":"","firstName":"Bienvenido","middleName":"","lastName":"Martínez-Navarro","suffix":""},{"id":286229077,"identity":"c748573c-ddbb-444c-a34d-136161b86419","order_by":3,"name":"Antonio Guerra-Merchán","email":"","orcid":"","institution":"University of Malaga: Universidad de Malaga","correspondingAuthor":false,"prefix":"","firstName":"Antonio","middleName":"","lastName":"Guerra-Merchán","suffix":""},{"id":286229078,"identity":"683c2830-805d-4ba5-af7d-0c5c8cbd0734","order_by":4,"name":"M. Dolores Rodríguez-Ruiz","email":"","orcid":"","institution":"University of Malaga: Universidad de Malaga","correspondingAuthor":false,"prefix":"","firstName":"M.","middleName":"Dolores","lastName":"Rodríguez-Ruiz","suffix":""},{"id":286229079,"identity":"f493bfb3-e614-4ce6-89ee-8fb3d17934c4","order_by":5,"name":"Guillermo Rodríguez-Gómez","email":"","orcid":"","institution":"Complutense University of Madrid: Universidad Complutense de Madrid","correspondingAuthor":false,"prefix":"","firstName":"Guillermo","middleName":"","lastName":"Rodríguez-Gómez","suffix":""},{"id":286229080,"identity":"a590dcc6-df97-41ad-a37b-5090f25dabae","order_by":6,"name":"Alejandro Granados","email":"","orcid":"","institution":"University of Malaga: Universidad de Malaga","correspondingAuthor":false,"prefix":"","firstName":"Alejandro","middleName":"","lastName":"Granados","suffix":""},{"id":286229081,"identity":"9206c99b-6c50-4df5-b5a1-cac72979b693","order_by":7,"name":"Laura León-Reina","email":"","orcid":"","institution":"University of Malaga: Universidad de Malaga","correspondingAuthor":false,"prefix":"","firstName":"Laura","middleName":"","lastName":"León-Reina","suffix":""},{"id":286229082,"identity":"d7c60052-f5f8-41e5-8fc6-9412a1ed03f2","order_by":8,"name":"M. Patrocinio Espigares","email":"","orcid":"","institution":"University of Malaga: Universidad de Malaga","correspondingAuthor":false,"prefix":"","firstName":"M.","middleName":"Patrocinio","lastName":"Espigares","suffix":""},{"id":286229083,"identity":"d3b48596-c513-4b8c-8782-3017f67a28ed","order_by":9,"name":"Sergio Ros-Montoya","email":"","orcid":"","institution":"University of Malaga: Universidad de Malaga","correspondingAuthor":false,"prefix":"","firstName":"Sergio","middleName":"","lastName":"Ros-Montoya","suffix":""},{"id":286229084,"identity":"d9a6665e-75f7-4641-b6bf-e50f54d6c80a","order_by":10,"name":"Paul Palmqvist","email":"data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAZAAAAAyAQMAAABI0h/eAAAABlBMVEX///8AAABVwtN+AAAACXBIWXMAAA7EAAAOxAGVKw4bAAAAi0lEQVRIiWNgGAWjYFACHsYHJGthNiBZC5sEaRrMpXuPVfPm2DHw8x8gUovlnHNpt3m3JTNIzkggUovBjRwzoBZmIINYh4G0FPNuq2ewP0+sw0BamHm3HWYwYCDWYUC/JEvO3XacR+IGsVqAIXbww9tt1XL8/UQ7DBopPESqR9IyCkbBKBgFowA3AABdlCPFN/cNpwAAAABJRU5ErkJggg==","orcid":"https://orcid.org/0000-0002-6630-6956","institution":"Universidad de Malaga","correspondingAuthor":true,"prefix":"","firstName":"Paul","middleName":"","lastName":"Palmqvist","suffix":""}],"badges":[],"createdAt":"2024-03-22 19:45:31","currentVersionCode":1,"declarations":"","doi":"10.21203/rs.3.rs-4151726/v1","doiUrl":"https://doi.org/10.21203/rs.3.rs-4151726/v1","draftVersion":[],"editorialEvents":[],"editorialNote":"","failedWorkflow":false,"files":[{"id":54073253,"identity":"0773c14d-f8a1-4b10-8875-61876384e2a2","added_by":"auto","created_at":"2024-04-04 07:59:36","extension":"jpg","order_by":1,"title":"Figure 1","display":"","copyAsset":false,"role":"figure","size":3652371,"visible":true,"origin":"","legend":"\u003cp\u003e\u003cstrong\u003eA:\u003c/strong\u003e High-scale geological context of the Baza Basin; the points mark the hot springs that are currently active. \u003cstrong\u003eB:\u003c/strong\u003e Geologic map of the Baza Basin. 1-Substrate of the External Zones; 2-Substrate of the Internal Zones; 3-Tortonian, marine facies; 4-Late Turolian-Pliocene, alluvial facies s.l.; 5-Pliocene-Lower Villafranchian, mainly lacustrine facies; 6-Early and Middle Villafranchian, distal fluvial facies (sandstones and lutites); 7-Early and Middle Villafranchian, marly-evaporitic lacustrine facies; 8-Late Villafranchian-Epivillafranchian, lacustrine and distal fluvial facies; 9-Middle and Late Pleistocene, proximal alluvial and locally lacustrine facies; 10-Glacis surface; 11-Holocene alluvial; 12-Holocene fluvial terraces; 13-Tectonic fractures (in discontinuous trace, those inferred from satellite images); 14-Hot springs currently active (codes match those of Table 2); 15-Recent travertine buildings; 16-Seismites; 17-Levels of dark lutites; 18-Cherts. Codes in red indicate the position of the stratigraphic series sampled (see Fig. 2). * Indicates the position of samples BNA-1, BNA-2, BNA-3, BNA-4, BNA-5, and BN-N at the northeastern of the locality of Benamaurel (Tables 3–4). \u003cstrong\u003eC: \u003c/strong\u003eGeographic and geologic context of the Orce-Venta Micena sector in the Baza Basin and lithostratigraphic model of the Orce subbasin with definition of tectono-sedimentary units (TSU) for the Baza Basin:1-Dolomites and limestones of the Subbetic domain (External Zones); 2-Alluvial red conglomerates; 3-Lacustrine yellow marls; 4-Lacustrine limestones; 5-Lacustrine marls, sands and gypsum; 6-Fluvial and lacustrine red and grey sands, as well as (calci)lutites; 7-Lacustrine marls and limestones; 8-Alluvial conglomerates; 9-Deltaic conglomerates and sands, as well as lacustrine marls; 10-Glacis surface; 11-Fluvial gravels and sands. Stratigraphic sections: Site codes and samples are depicted in Figure 2. The equivalence between the codes that appear in Figures 1B–C is the following:1–1,\u003cstrong\u003e \u003c/strong\u003e4–2 \u0026amp; 8,\u003cstrong\u003e \u003c/strong\u003e5–3 \u0026amp; 4,\u003cstrong\u003e \u003c/strong\u003e6–5,\u003cstrong\u003e \u003c/strong\u003e7–5\u003cstrong\u003e, \u003c/strong\u003e8–6 \u0026amp; 7,\u003cstrong\u003e \u003c/strong\u003e9–9\u003cstrong\u003e, \u003c/strong\u003e10–10, and\u003cstrong\u003e \u003c/strong\u003e11–11.\u003c/p\u003e","description":"","filename":"Figure1.jpg","url":"https://assets-eu.researchsquare.com/files/rs-4151726/v1/adb317678f995aa807d50a6b.jpg"},{"id":54073255,"identity":"794f8c41-ab57-4cff-92b8-f2225817bd7a","added_by":"auto","created_at":"2024-04-04 07:59:37","extension":"jpg","order_by":2,"title":"Figure 2","display":"","copyAsset":false,"role":"figure","size":3997021,"visible":true,"origin":"","legend":"\u003cp\u003eStratigraphic profiles of the Baza Basin. The codes with the position of these series are shown in Figures 1B–C.\u003c/p\u003e","description":"","filename":"Figure2.jpg","url":"https://assets-eu.researchsquare.com/files/rs-4151726/v1/403f97fa73e84d1b80c9812d.jpg"},{"id":54073258,"identity":"a6f3ae12-a949-4444-a5f6-f86d504bebfd","added_by":"auto","created_at":"2024-04-04 07:59:37","extension":"jpg","order_by":3,"title":"Figure 3","display":"","copyAsset":false,"role":"figure","size":12536220,"visible":true,"origin":"","legend":"\u003cp\u003eField images of cherts and other facies linked to hydrothermal scenarios and associated to tectonic fractures. \u003cstrong\u003eA:\u003c/strong\u003e light colored chert deposits located at NW of the Orce town (sample SX-A, Fig. 1C); \u003cstrong\u003eB:\u003c/strong\u003e detail of white cherts located close to the FN1 paleontological site (sample FN1A, Fig. 2); \u003cstrong\u003eC:\u003c/strong\u003e thin level of grey to blue cherts (just at the top of the hammer), located at south of the Galera town (sample GA-6, Fig 2); note the presence of deformed layers at the bottom of this image. \u003cstrong\u003eD:\u003c/strong\u003e dark level located at the BZ-1 Pliocene site (sample BZ-1A, Fig. 2). \u003cstrong\u003eE:\u003c/strong\u003e travertines (sample TE100, Fig. 1C) located west of Orce. \u003cstrong\u003eF:\u003c/strong\u003e detail of these travertine facies. \u003cstrong\u003eG:\u003c/strong\u003e NE-SW oriented tectonic fault present at the south of Galera (marked with an orange arrow), which affects the chert levels; the fault plane appears to be filled with gypsum ore. \u003cstrong\u003eH:\u003c/strong\u003e mineral association between gypsum crystals (marked with an orange circle) and cherts, near the locality of Galera. \u003cstrong\u003eI: \u003c/strong\u003e~3 m wide chert dyke (the limits of the dyke are marked with orange arrows) with a NE-SW orientation, which coincides with a tectonic fracture near Galera (Fig. 1C). \u003cstrong\u003eJ: \u003c/strong\u003edetail of the N105ºE tectonic contact between a chert layer and a deposit of calciarenites (marked with an orange arrow), which appears close to the paleontological site of FN1 (Fig. 1C).\u003c/p\u003e","description":"","filename":"Figure3.jpg","url":"https://assets-eu.researchsquare.com/files/rs-4151726/v1/d83dc3b345dbabca00372acc.jpg"},{"id":54073264,"identity":"5ebe1c35-9b8f-45c6-83b2-5c6057d0b63c","added_by":"auto","created_at":"2024-04-04 07:59:37","extension":"jpg","order_by":4,"title":"Figure 4","display":"","copyAsset":false,"role":"figure","size":596793,"visible":true,"origin":"","legend":"\u003cp\u003eAl-Fe-Mn diagram applied to all samples analyzed in this study. Green area indicates a hydrothermal origin (Adachi et al., 1986; Yamamoto, 1987).\u003c/p\u003e","description":"","filename":"Figure4.jpg","url":"https://assets-eu.researchsquare.com/files/rs-4151726/v1/f40487668c07be0d687376a7.jpg"},{"id":54073259,"identity":"5d04897f-809a-4130-be28-093a9b72b6a9","added_by":"auto","created_at":"2024-04-04 07:59:37","extension":"jpg","order_by":5,"title":"Figure 5","display":"","copyAsset":false,"role":"figure","size":1200833,"visible":true,"origin":"","legend":"\u003cp\u003eX-ray diffraction diagrams of the studied cherts. Abbreviations for minerals follow Warr (2021). For more information, see Table 6.\u003c/p\u003e","description":"","filename":"Figure5.jpg","url":"https://assets-eu.researchsquare.com/files/rs-4151726/v1/c34116a08cccedf614d32268.jpg"},{"id":54073261,"identity":"b024b5a9-d2a9-42c3-a740-01295fa49ee8","added_by":"auto","created_at":"2024-04-04 07:59:37","extension":"jpg","order_by":6,"title":"Figure 6","display":"","copyAsset":false,"role":"figure","size":229866,"visible":true,"origin":"","legend":"\u003cp\u003eSpatio-temporal position of the samples with a hydrothermal origin and of those hot springs currently active in the Baza Basin. The X-axis indicates a SW-NE line along the Baza Basin, between the localities of Zújar and Cañada del Negro (NE of Orce), with a length of ~50 km. Key for localities: ZU-Zújar, BZ-Baza, BN-Benamaurel, GA-Galera, HU-Huéscar, OR-Orce, CN-Cañada del Negro. Key of chronological periods (Y-axis): R-Ruscinian, EV-Early Villafranchian, MV-Middle Villafranchian, LV-Late Villafranchian, E-Epivillafranchian, G-Galerian, A-Aurelian. Red points show the position of the samples linked to a hydrothermal context. Blue points show the position of the hot springs currently active in the Baza Basin (code numbers are similar to those of Table 2).\u003c/p\u003e","description":"","filename":"Figure6.jpg","url":"https://assets-eu.researchsquare.com/files/rs-4151726/v1/14924051a43fd833bb390c34.jpg"},{"id":54073265,"identity":"8f0698dc-c5e8-45d8-8565-330287a010c8","added_by":"auto","created_at":"2024-04-04 07:59:37","extension":"jpg","order_by":7,"title":"Figure 7","display":"","copyAsset":false,"role":"figure","size":1994282,"visible":true,"origin":"","legend":"\u003cp\u003ePaleoecological reconstruction of the landscape in the vicinity of a lacustrine system of the Baza Basin linked to a thermal upwelling. \u003cstrong\u003eA:\u003c/strong\u003e Highstand level in the lacustrine system. \u003cstrong\u003eB: \u003c/strong\u003eLowstand level in the lacustrine system. The trace of the fault positioned to the left indicates the position of a local hot spring. Drawing by Sandra Ramírez Cherbuy - InkScience.\u003c/p\u003e","description":"","filename":"Figure7.jpg","url":"https://assets-eu.researchsquare.com/files/rs-4151726/v1/44b679f22a82561f6f84b1c5.jpg"},{"id":54073262,"identity":"d67e837e-2561-4e29-a70a-9310e5ec3248","added_by":"auto","created_at":"2024-04-04 07:59:37","extension":"jpg","order_by":8,"title":"Figure 8","display":"","copyAsset":false,"role":"figure","size":1075052,"visible":true,"origin":"","legend":"\u003cp\u003e\u003cstrong\u003eA: \u003c/strong\u003eLeast-squares relationship between the logarithms of annual precipitation (mm) and herbivore prey biomass (kg/km\u003csup\u003e2\u003c/sup\u003e) in 54 National Parks and Game Reserves of Africa [data on mean annual rainfall from Fick and Hijmans (2017); data on prey biomass from Bell (1981) and Hatton et al. (2015)]. The values estimated for the paleontological site of Venta Micena (VM) and the archaeological sites of Barranco León (BL) and Fuente Nueva-3 (FN-3) in the Baza Basin, not used for deriving the equation, are also shown [rainfall estimates based on Blain et al. (2016) for BL and FN3, and Palmqvist et al. (2022c) for VM; prey biomass estimates taken from Rodríguez-Gómez et al. (2022)]. \u003cstrong\u003eB:\u003c/strong\u003e Whiskler box-plots for the unstandardized residuals of the regression equation of A, differentiating between sediment types and between localities with or without thermal activity.\u003c/p\u003e","description":"","filename":"Figure8.jpg","url":"https://assets-eu.researchsquare.com/files/rs-4151726/v1/0337a54774ec7dcbd433c8d2.jpg"},{"id":54073263,"identity":"99caf96f-333b-41f2-9082-0154eb8d3435","added_by":"auto","created_at":"2024-04-04 07:59:37","extension":"jpg","order_by":9,"title":"Figure 9","display":"","copyAsset":false,"role":"figure","size":2135081,"visible":true,"origin":"","legend":"\u003cp\u003eMap of Africa and Western Eurasia showing the position of selected archaeological and paleontological sites, as well as the geologically active areas. Site codes and references: 1: Ain Maarouf (Geraads et al., 1992); 2: Djebel Irhoud (Hublin et al., 2017); 3: Tighenif (Sahnouni and Van der Made, 2009); 4: N’Gaous and Ket Sefiane (Sala et al., 2016); 5: Ain Hanech, El Kherba and Ain Boucherit (Sahnouni et al., 2018); 6: Oued Sarrat (Martínez-Navarro et al., 2014); 7: Toros Menalla (Brunet et al., 2002); 8: Qesem Cave (Barkai et al., 2003); 9: Gesher B. Ya’akov and Ma’ayan Barukh (Goren-Inbar et al., 2000; Ronen et al., 1980); 10: Ubeidiya, Evron Quarry, Revadim Quarry, and Bizat Ruhama (Tchernov et al., 1987; Tchernov et al., 1994; Martínez-Navarro et al., 2012; Yeshurun et al., 2011; Marder et al., 2007); 11: Latamne (Guérin et al., 1993); 12: Ti´s Al Gahdah (Stimpson et al., 2016); 13: Buia and Engel Ela-Ramud (Abbate et al., 1998; Martínez-Navarro et al., 2016); 14: Hadar and Dikkita-Dikia (Johanson et al., 1978; Alemseged et al., 2006); 15: Gona I and II (Semaw et al., 1997); 16: Middle-Awash, Aramis I and II, Maka, and Bouri (White et al., 1993; White et al., 1994; White et al., 2003; Asfaw et al., 2002); 17: Fejej (Barsky et al., 2011); 18: Gadeb (Williams et al., 1979); 19: Melka-Kunture (Morgan et al., 2012); 20: Asbole and Bodo (Geraads et al., 2004; Conroy et al., 1978); 21: Konso-Gardula (Asfaw et al., 1992); 22: Shungura (Boaz and Howell, 1977; Howell et al., 1987); 23: Woranso Mille (Haile-Selassie et al., 2010); 24: Koobi-Fora (Bunn, 1981; Lepre et al., 2007; Lepre and Kent, 2015); 25: Lokalalei and Lomekwi (Harmand et al., 2015); 26: Ileret (Jungers et al., 2015); 27: Chesowanja (Gowlett et al., 1981); 28: Kilombe and Kapturin (Gowlett, 2005); 29: Kanjera (Plummer and Bishop, 2016); 30: Kariandusi (Shipton, 2011); 31: Kanapoi and Lothagam (Leakey et al., 1995; Kissel and Hawks, 2015); 32: Lukeino-Tugen Hills (Sawada et al., 2002); 33: Kantis (Mbua et al., 2016); 34: Olorgesailie (Potts, 1994); 35: Nyabusosi (Texier, 1995); 36: Peninj (De la Torre et al., 2003); 37: Olduvai (Leakey and Leakey, 1964); 38: Laetoli (Magori and Day, 1983); 39: Kalombo Falls (Duller et al., 2015); 40: Uraha Hill and Chiwondo (Bromage et al., 1995); 41: Sterkfontein, Swartkrans, Drimden, Makapansgat, Malapa, Haasgat Cave, Kromdraai Cave, Cooper’s Cave, and Gladysvale Cave (Herries, 2011; Herries and Saw, 2011; Herries and Adams, 2013; Herries et al., 2014; Herries et al., 2018; Latham and Herries, 2004; Dirks et al., 2010; Kuman and Clarke, 2000; Kuman, 2007; De Ruiter et al., 2009); 42: Taung (McKee, 1993); 43: Wonderwerk Cave (Chazan et al., 2008); 44: Elandsfontein (Braun et al., 2013); 45: Aroeira (Daura et al., 2017); 46: Atapuerca (Carbonell et al., 2008; Bermúdez de Castro et al., 1997; Parés and Pérez-González, 1999); 47: Valparadis (Martínez et al., 2013); 48: Alto de las Picarazas (Gabarda et al., 2016); 49: Orce, Cúllar and Solana del Zamborino (Martínez-Navarro et al., 1997; Oms et al., 2000a; Oms et al., 2000b; Toro-Moyano et al., 2013; Espigares et al. 2013, Torrente Casado, 2010; Álvarez-Posada et al., 2017); 50: Cueva Victoria (Gibert et al., 2016); 51: Bois-de-Riquet and Vallonet Cave (Bourguignon et al., 2016; Michel et al., 2017); 52: Tautavel-Arago (Byrne, 2004; Barsky and de Lumley, 2010); 53: Pont-de-Lavaud (Bahain et al., 2007; Lombera-Hermida et al., 2016); 54: Lunery-Rosières (Despriée et al., 2009); 55: Happisburgh (Parfitt et al., 2010); 56: Pakefield (Parfitt et al., 2005); 57: Swanscombe (Lister, 1986); 58: Pirro Nord (López-García et al., 2015); 59: Ca’Belvedere di Monte Poggiolo (Bahain et al., 2007); 60: Isernia la Pineta (Coltorti et al., 2005; Bahain et al., 2007); 61: Ciampate di Diavolo (Mietto et al., 2003); 62: Untermassfeld (Baales, 2014); 63: Petralona Cave (Poulianos, 1984); 64: Kozarnika Cave I and II (Sirakov et al., 2010); 65: Dmanisi and Akhalkalaki (Lordkipanidze et al., 2007; Vekua,1986); 66: Muhkai II (Amirkhanov et al., 2016; Michel et al., 2017); 67: Kermek, Rodniki, Bogatyri-Sinyaya, Balka, and Tsymbal (Shchelinsky et al., 2010; Michel et al., 2017).\u003c/p\u003e","description":"","filename":"Figure9.jpg","url":"https://assets-eu.researchsquare.com/files/rs-4151726/v1/ff58015b477111354e3c7362.jpg"},{"id":54073864,"identity":"42610d0f-4af6-4e51-8105-4af895c31a06","added_by":"auto","created_at":"2024-04-04 08:07:45","extension":"pdf","order_by":0,"title":"","display":"","copyAsset":false,"role":"manuscript-pdf","size":1986429,"visible":true,"origin":"","legend":"","description":"","filename":"manuscript.pdf","url":"https://assets-eu.researchsquare.com/files/rs-4151726/v1/f091bd02-02d7-4715-a410-c9b81f78d9da.pdf"},{"id":54073251,"identity":"f83e6315-03c0-45d8-a238-6ed516b3258b","added_by":"auto","created_at":"2024-04-04 07:59:36","extension":"docx","order_by":1,"title":"","display":"","copyAsset":false,"role":"supplement","size":22916,"visible":true,"origin":"","legend":"","description":"","filename":"Table1.docx","url":"https://assets-eu.researchsquare.com/files/rs-4151726/v1/63279de89224602d3c21cb56.docx"},{"id":54073848,"identity":"95f997c2-6437-4bbf-9b0d-f9b15dc63886","added_by":"auto","created_at":"2024-04-04 08:07:36","extension":"docx","order_by":2,"title":"","display":"","copyAsset":false,"role":"supplement","size":25085,"visible":true,"origin":"","legend":"","description":"","filename":"Table2.docx","url":"https://assets-eu.researchsquare.com/files/rs-4151726/v1/f1b2b5c02b0dc832868e18f1.docx"},{"id":54073257,"identity":"7d169cc0-2e8a-41aa-a3d4-33e451c552ae","added_by":"auto","created_at":"2024-04-04 07:59:37","extension":"docx","order_by":3,"title":"","display":"","copyAsset":false,"role":"supplement","size":45781,"visible":true,"origin":"","legend":"","description":"","filename":"Table3.docx","url":"https://assets-eu.researchsquare.com/files/rs-4151726/v1/8a755877f94ba3aa9c2fd217.docx"},{"id":54073254,"identity":"22867bbe-ea77-4e3f-83eb-2b1148887219","added_by":"auto","created_at":"2024-04-04 07:59:36","extension":"docx","order_by":4,"title":"","display":"","copyAsset":false,"role":"supplement","size":41371,"visible":true,"origin":"","legend":"","description":"","filename":"Table4.docx","url":"https://assets-eu.researchsquare.com/files/rs-4151726/v1/bbc8028009d9c92ed198e088.docx"},{"id":54073849,"identity":"34997089-cc60-41e8-a002-a5ce1c6f2812","added_by":"auto","created_at":"2024-04-04 08:07:37","extension":"docx","order_by":5,"title":"","display":"","copyAsset":false,"role":"supplement","size":27851,"visible":true,"origin":"","legend":"","description":"","filename":"Table5.docx","url":"https://assets-eu.researchsquare.com/files/rs-4151726/v1/5d85cbb39bbaec89866f2d8d.docx"},{"id":54073256,"identity":"232f669c-9156-4967-9449-96aeb04a223f","added_by":"auto","created_at":"2024-04-04 07:59:37","extension":"docx","order_by":6,"title":"","display":"","copyAsset":false,"role":"supplement","size":19957,"visible":true,"origin":"","legend":"","description":"","filename":"Table6.docx","url":"https://assets-eu.researchsquare.com/files/rs-4151726/v1/6e65a8c849476ba0625e8c16.docx"}],"financialInterests":"","formattedTitle":"Plio-Pleistocene hydrothermal events of the Baza Basin (Betic Cordillera, SE Spain) and their paleoecological implications","fulltext":[{"header":"1 Introduction and background","content":"\u003cp\u003eThe Baza Basin lies in the northeastern sector of the Guadix-Baza Depression (Granada, SE Spain; Fig. \u003cspan class=\"InternalRef\"\u003e1\u003c/span\u003eA), which extends over an area of ~\u0026thinsp;4000 km\u003csup\u003e2\u003c/sup\u003e (Palmqvist et al., \u003cspan class=\"CitationRef\"\u003e2022a\u003c/span\u003e) and is surrounded by the highest reliefs of the Alpine Betic cordillera. This inland basin preserves a unique Plio-Pleistocene sedimentary record of high preservational completeness, which is composed of fluvio-lacustrine deposits that preserve many paleontological and archaeological localities distributed throughout the whole sedimentary depression, and especially in the Baza Basin (Mart\u0026iacute;nez-Navarro, \u003cspan class=\"CitationRef\"\u003e1991\u003c/span\u003e; Palmqvist et al., \u003cspan class=\"CitationRef\"\u003e1996\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2011\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2022a\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2022b\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2023\u003c/span\u003e; Arribas and Palmqvist, \u003cspan class=\"CitationRef\"\u003e1998\u003c/span\u003e; Palmqvist and Arribas, \u003cspan class=\"CitationRef\"\u003e2001\u003c/span\u003e; Espigares et al., \u003cspan class=\"CitationRef\"\u003e2013\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2019\u003c/span\u003e; Toro-Moyano et al., \u003cspan class=\"CitationRef\"\u003e2013\u003c/span\u003e; Rodr\u0026iacute;guez-G\u0026oacute;mez et al., \u003cspan class=\"CitationRef\"\u003e2016\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2017\u003c/span\u003e; Ros-Montoya et al., \u003cspan class=\"CitationRef\"\u003e2017\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2018\u003c/span\u003e; Mart\u0026iacute;nez-Navarro et al., \u003cspan class=\"CitationRef\"\u003e2018\u003c/span\u003e; Table \u003cspan class=\"InternalRef\"\u003e1\u003c/span\u003e). The presence of Plio-Pleistocene hydrothermal events in the Baza Basin and their paleoecological consequences has been documented since the end of the 20th century by means of sedimentological, tectonic, and mineralogical data (e.g., Sebasti\u0026aacute;n Pardo, \u003cspan class=\"CitationRef\"\u003e1979\u003c/span\u003e; Garc\u0026iacute;a-Aguilar, \u003cspan class=\"CitationRef\"\u003e1997\u003c/span\u003e; Garc\u0026iacute;a-Aguilar et al., \u003cspan class=\"CitationRef\"\u003e2014\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2015\u003c/span\u003e; S\u0026aacute;nchez-Roa et al., \u003cspan class=\"CitationRef\"\u003e2016\u003c/span\u003e).\u003c/p\u003e\n\u003cdiv id=\"Sec2\" class=\"Section2\"\u003e\n \u003ch2\u003e1.1 Geological and paleontological setting\u003c/h2\u003e\n \u003cp\u003eAccording to the classification of Miall (\u003cspan class=\"CitationRef\"\u003e1984\u003c/span\u003e), the Baza Basin is an intramountainous basin with tectonic subsidence and continental sedimentation, features inherited from an ancient marine basin formed during the Late Miocene in the interior of the Betic mountain range (Sanz de Galdeano and Vera, 1992; Platt et al., \u003cspan class=\"CitationRef\"\u003e2013\u003c/span\u003e). The transition from a marine to a continental sedimentary regime took place 7.5\u0026ndash;7.0 Ma ago, being the result of major regional tectonic movements that caused the isostatic uplifting of the basin (average uplift rate of ~\u0026thinsp;200 m/Ma, as estimated from Late Neogene coastal marine conglomerates and coral reefs; Braga et al., \u003cspan class=\"CitationRef\"\u003e2003\u003c/span\u003e), which resulted in its disconnection from the Mediterranean Sea (Soria, \u003cspan class=\"CitationRef\"\u003e1993\u003c/span\u003e; Garc\u0026iacute;a-Aguilar, \u003cspan class=\"CitationRef\"\u003e1997\u003c/span\u003e; Play\u0026aacute; et al., 2000; Guerra-Merch\u0026aacute;n et al., \u003cspan class=\"CitationRef\"\u003e2014\u003c/span\u003e). The Baza Basin is bounded to the north by reliefs of Mesozoic age, mostly carbonates (Thetys type), which belong to the External Zones of the Betic Cordillera. To the south, it is surrounded by reliefs composed of sedimentary and metamorphic rocks of Paleozoic-Triassic age, which belong to the Internal Zones. To the west, it is connected with the Neogene Guadix Basin and to the southeast with the Neogene Almanzora Corridor Basin (Fig. \u003cspan class=\"InternalRef\"\u003e1\u003c/span\u003eB).\u003c/p\u003e\n \u003cp\u003eThe Plio-Pleistocene stratigraphic model of the Baza Basin shows three major stages (Fig. \u003cspan class=\"InternalRef\"\u003e1\u003c/span\u003eC), which are represented by tectono-sedimentary units (TSU) according to Meg\u0026iacute;as (\u003cspan class=\"CitationRef\"\u003e1982\u003c/span\u003e) (see also Guerra-Merch\u0026aacute;n, \u003cspan class=\"CitationRef\"\u003e1992\u003c/span\u003e; Soria, \u003cspan class=\"CitationRef\"\u003e1993\u003c/span\u003e; Garc\u0026iacute;a-Aguilar, \u003cspan class=\"CitationRef\"\u003e1997\u003c/span\u003e; Garc\u0026iacute;a-Aguilar and Mart\u0026iacute;n, \u003cspan class=\"CitationRef\"\u003e2000\u003c/span\u003e; Garc\u0026iacute;a-Aguilar and Palmqvist, \u003cspan class=\"CitationRef\"\u003e2011\u003c/span\u003e). The three TSUs are bounded by sedimentary hiatuses and/or unconformities that represent stages in which the basin was subject to intense tectonic activity, showing little or no sedimentation. These units have been dated using biochronological (e.g., Alberdi and Bonadonna, \u003cspan class=\"CitationRef\"\u003e1989\u003c/span\u003e; Mart\u0026iacute;nez-Navarro, \u003cspan class=\"CitationRef\"\u003e1991\u003c/span\u003e; Ses\u0026eacute;, \u003cspan class=\"CitationRef\"\u003e1991\u003c/span\u003e; Guerra-Merch\u0026aacute;n, \u003cspan class=\"CitationRef\"\u003e1992\u003c/span\u003e; Oms et al., \u003cspan class=\"CitationRef\"\u003e2000a\u003c/span\u003e; Ses\u0026eacute;, \u003cspan class=\"CitationRef\"\u003e2006\u003c/span\u003e; Gibert and Gibert, \u003cspan class=\"CitationRef\"\u003e2003\u003c/span\u003e; Ros-Montoya, \u003cspan class=\"CitationRef\"\u003e2010\u003c/span\u003e; Maldonado-Garrido et al., \u003cspan class=\"CitationRef\"\u003e2017\u003c/span\u003e) and paleomagnetic data (e.g., Garc\u0026eacute;s, \u003cspan class=\"CitationRef\"\u003e1994\u003c/span\u003e; Oms et al., \u003cspan class=\"CitationRef\"\u003e1994\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e1996\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e1999\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2000b\u003c/span\u003e; Gibert et al., \u003cspan class=\"CitationRef\"\u003e2006\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2007\u003c/span\u003e; Scott et al., \u003cspan class=\"CitationRef\"\u003e2007\u003c/span\u003e; Oms et al., \u003cspan class=\"CitationRef\"\u003e2011\u003c/span\u003e; \u0026Aacute;lvarez et al., \u003cspan class=\"CitationRef\"\u003e2015\u003c/span\u003e; \u0026Aacute;lvarez-Posada et al., \u003cspan class=\"CitationRef\"\u003e2017\u003c/span\u003e).\u003c/p\u003e\n \u003cp\u003eThe first TSU, which is of Pliocene age, shows a sediment thickness of ~\u0026thinsp;150 m. It is characterized by the alternation of marls/calcilutites/lutites and limestones deposited in lacustrine systems, which change towards the edges of the basin to sands and conglomerates of alluvial origin. Forest ecosystems were formed on the margins of the lake as inferred for the Ruscinian (~\u0026thinsp;4.5 Ma) Baza-1 paleontological site (Ros-Montoya et al., \u003cspan class=\"CitationRef\"\u003e2017\u003c/span\u003e). Tectonic activity during this stage induced the formation of many soft-sediment deformation structures (i.e., paleo-seismites), which are mainly located at the marginal sites of a paleo-lake (Alfaro et al., \u003cspan class=\"CitationRef\"\u003e1997\u003c/span\u003e) and possibly resulted in deposits with hydrothermal affinities (Garc\u0026iacute;a-Aguilar et al., \u003cspan class=\"CitationRef\"\u003e2014\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2015\u003c/span\u003e). Climate was warm and humid during this stage, which relates to an environment typical of modern subtropical or rainy tropical climates (Foucault and M\u0026eacute;li\u0026egrave;res, \u003cspan class=\"CitationRef\"\u003e2000\u003c/span\u003e; Haywood and Vald\u0026eacute;s, \u003cspan class=\"CitationRef\"\u003e2004\u003c/span\u003e).\u003c/p\u003e\n \u003cp\u003eThe second TSU has a latest Pliocene to Middle Pleistocene age and is the most complex, both lithologically and tectonically. During this stage, a great paleogeographic change took place in the Baza Basin, which involved an ~\u0026thinsp;E-W expansion linked to an NNW-SSE fracture system. This in turn resulted in intense subsidence and the deposition of sediments with a minimum thickness of ~\u0026thinsp;280 m in the central sector of the basin. However, geophysical data (Sanz de Galdeano et al., 2007) indicate that sedimentary thickness may be much greater than the one visible. These sediments, contributed by fluvial systems from the adjacent Guadix Basin, filled a large lake of marly-evaporitic sedimentation with an outer rim of detrital inputs (Vera, \u003cspan class=\"CitationRef\"\u003e1970\u003c/span\u003e; Garc\u0026iacute;a-Aguilar and Mart\u0026iacute;n, \u003cspan class=\"CitationRef\"\u003e2000\u003c/span\u003e; Garc\u0026iacute;a-Aguilar and Palmqvist, \u003cspan class=\"CitationRef\"\u003e2011\u003c/span\u003e). Tectonic activity during this stage resulted in synsedimentary normal faults, tilting, local unconformities, and seismites (Vera et al., \u003cspan class=\"CitationRef\"\u003e1984\u003c/span\u003e; Alfaro et al., \u003cspan class=\"CitationRef\"\u003e1997\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2000\u003c/span\u003e). The lower section of this unit shows selenitic and microcrystalline gypsum levels of centimeter thickness, which alternate with layers of light-colored marls towards the basin center or with sandstones towards the edges (Garc\u0026iacute;a-Aguilar et al., \u003cspan class=\"CitationRef\"\u003e2013\u003c/span\u003e). Climate at this stage was semi-arid, with cyclic oscillations within the Bond frequency timing band (Hagelberg et al., \u003cspan class=\"CitationRef\"\u003e1994\u003c/span\u003e; Bond et al., \u003cspan class=\"CitationRef\"\u003e1997\u003c/span\u003e; Garc\u0026iacute;a-Aguilar et al., \u003cspan class=\"CitationRef\"\u003e2013\u003c/span\u003e), which generated alternating periods of inundation and desiccation of the lake linked to the deposition of marls or sandstones and gypsum, respectively.\u003c/p\u003e\n \u003cp\u003eThe upper section of this TSU outcrops mainly in the Orce area, a sub-basin in the NE sector of the Baza Basin, which has an extent of ~\u0026thinsp;170 km\u003csup\u003e2\u003c/sup\u003e (Figs. \u003cspan class=\"InternalRef\"\u003e1\u003c/span\u003eB\u0026ndash;C) and shows extensive outcrops\u0026thinsp;~\u0026thinsp;35 m thick that correspond to the Early to Middle Pleistocene. The Orce sub-basin preserves a unique fossil record of terrestrial vertebrates, including two archaeological sites dated to ~\u0026thinsp;1.4 Ma (see review in Palmqvist et al., \u003cspan class=\"CitationRef\"\u003e2016\u003c/span\u003e) that mark the earliest presence of hominins in Western Europe: Barranco Le\u0026oacute;n (BL) and Fuente Nueva-3 (FN3). BL and FN3 preserve huge assemblages of skeletal remains of large mammals and Oldowan tools, which are linked through the presence of anthropogenic cut and percussion marks, as well as a human deciduous tooth in the case of BL, which is at this moment the oldest hominin fossil found in Western Europe (Mart\u0026iacute;nez-Navarro et al., \u003cspan class=\"CitationRef\"\u003e1997\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2014\u003c/span\u003e; Oms et al., \u003cspan class=\"CitationRef\"\u003e2000a\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2000b\u003c/span\u003e; Espigares et al., \u003cspan class=\"CitationRef\"\u003e2013\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2019\u003c/span\u003e; Toro-Moyano et al., \u003cspan class=\"CitationRef\"\u003e2013\u003c/span\u003e; Titton et al., \u003cspan class=\"CitationRef\"\u003e2021\u003c/span\u003e; Yravedra et al., \u003cspan class=\"CitationRef\"\u003e2021\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2022\u003c/span\u003e; Palmqvist et al., \u003cspan class=\"CitationRef\"\u003e2023\u003c/span\u003e). Moreover, the Venta Micena site (VM, 1.6\u0026ndash;1.5 Ma), which is slightly older than BL/FN3 and has been interpreted as a denning area of the giant hyena \u003cem\u003ePachycrocuta brevirostris\u003c/em\u003e (Arribas and Palmqvist, \u003cspan class=\"CitationRef\"\u003e1998\u003c/span\u003e; Palmqvist and Arribas, \u003cspan class=\"CitationRef\"\u003e2001\u003c/span\u003e; Palmqvist et al., \u003cspan class=\"CitationRef\"\u003e2022b\u003c/span\u003e), provides an excellent record of the large mammal community that preceded the hominin dispersal in the basin (Mart\u0026iacute;nez-Navarro, \u003cspan class=\"CitationRef\"\u003e1991\u003c/span\u003e; Mendoza et al., \u003cspan class=\"CitationRef\"\u003e2005\u003c/span\u003e; Rodr\u0026iacute;guez-G\u0026oacute;mez et al., \u003cspan class=\"CitationRef\"\u003e2017\u003c/span\u003e). The varied lithological composition of the Orce area, with alternations of fluvial red detrital facies and lacustrine carbonate-marly facies that locally show gypsum deposits (restricted to the center of the lacustrine system in this area) and cherts, points to a paleogeographic model of fluvio-lacustrine sedimentation highly variable in time and space. This resulted in open plains with forest patches in the emerged areas surrounding the fluvio-lacustrine system (Mendoza et al., \u003cspan class=\"CitationRef\"\u003e2005\u003c/span\u003e; Scott et al., \u003cspan class=\"CitationRef\"\u003e2007\u003c/span\u003e; Garc\u0026iacute;a-Aguilar and Palmqvist, \u003cspan class=\"CitationRef\"\u003e2011\u003c/span\u003e; Palmqvist et al., \u003cspan class=\"CitationRef\"\u003e2022a\u003c/span\u003e).\u003c/p\u003e\n \u003cp\u003eThe third TSU of the Baza Basin has a Middle Pleistocene to Late Pleistocene age, showing a maximum thickness of 70 m. During this stage, the activity of major faults (Baza and Galera) resulted in some tectonic blocks that developed an effective subsidence, which allowed a continuous sedimentation. This TSU is composed of conglomerates and sands towards the basin edges, which change to lutites, marls and marly limestones deposited in shallow lacustrine environments towards the basin center. In the upper part of this unit and at the basin edges, alluvial facies (mainly red conglomerates and sands) prograde over lacustrine facies. This TSU is represented in several sectors of the basin perimeter (Figs. \u003cspan class=\"InternalRef\"\u003e1\u003c/span\u003eB\u0026ndash;C) and the most significant paleontological sites are Hu\u0026eacute;scar-1 and C\u0026uacute;llar-Baza-1 (Ruiz Bustos, 1976; Pe\u0026ntilde;a et al., \u003cspan class=\"CitationRef\"\u003e1977\u003c/span\u003e; Mazo et al., \u003cspan class=\"CitationRef\"\u003e1985\u003c/span\u003e; Alberdi and Bonadonna, \u003cspan class=\"CitationRef\"\u003e1989\u003c/span\u003e; Agust\u0026iacute; et al., \u003cspan class=\"CitationRef\"\u003e2000\u003c/span\u003e; Torrente Casado, \u003cspan class=\"CitationRef\"\u003e2010\u003c/span\u003e; Ros-Montoya et al., \u003cspan class=\"CitationRef\"\u003e2018\u003c/span\u003e).\u003c/p\u003e\n \u003cp\u003eFinally, during the late Middle Pleistocene (~\u0026thinsp;400 ka: Garc\u0026iacute;a-Tortosa et al., \u003cspan class=\"CitationRef\"\u003e2008a\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003eb\u003c/span\u003e) or in Late Pleistocene times (100\u0026ndash;17 ka: Viseras and Fern\u0026aacute;ndez, \u003cspan class=\"CitationRef\"\u003e1992\u003c/span\u003e; Calvache and Viseras, \u003cspan class=\"CitationRef\"\u003e1997\u003c/span\u003e; 42 ka: Aza\u0026ntilde;\u0026oacute;n et al., \u003cspan class=\"CitationRef\"\u003e2006\u003c/span\u003e), the hydrographic network of the sedimentary depression was captured by the Guadiana Menor River, a tributary of the Guadalquivir River. This resulted in the transit from an endorheic regime to an exorheic one, leading to a stage in which erosion dominated over sedimentation. As a result, a badlands landscape predominates now in the region, linked to a mean annual precipitation (MAP) of ~\u0026thinsp;300 mm in the innermost part of the sedimentary depression (\u003cspan class=\"ExternalRef\"\u003e\u003cspan class=\"RefSource\"\u003ehttps://es.climate-data.org\u003c/span\u003e\u003c/span\u003e). However, climate was more humid during the late Early Pleistocene, with an estimated MAP of ~\u0026thinsp;700 mm in BL/FN3 and ~\u0026thinsp;850 mm in VM (Palmqvist et al., \u003cspan class=\"CitationRef\"\u003e2022c\u003c/span\u003e).\u003c/p\u003e\n\u003c/div\u003e\n\u003cdiv id=\"Sec3\" class=\"Section2\"\u003e\n \u003ch2\u003e1.2 Hot springs currently active in the Baza Basin\u003c/h2\u003e\n \u003cp\u003eTectonic dynamism resulted in hot springs, several of which are currently active in the Baza Basin and other nearby areas (Cruz-Sanjuli\u0026aacute;n et al., \u003cspan class=\"CitationRef\"\u003e1972\u003c/span\u003e; Cruz-Sanjuli\u0026aacute;n and Garc\u0026iacute;a-Rosell, \u003cspan class=\"CitationRef\"\u003e1972\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e1975\u003c/span\u003e; Diputaci\u0026oacute;n de Granada-ITGE, \u003cspan class=\"CitationRef\"\u003e1990\u003c/span\u003e; Figs. \u003cspan class=\"InternalRef\"\u003e1\u003c/span\u003eA\u0026ndash;B, and Table \u003cspan class=\"InternalRef\"\u003e2\u003c/span\u003e). There are twelve hot springs in the Baza Basin and four in the Guadix Basin, all of them associated with regional faults. In the case of the hot springs of the Baza Basin, their water output temperatures range between 18\u0026ordm; C and 22\u0026ordm; C, except in the case of the \u0026lsquo;Ba\u0026ntilde;os de Z\u0026uacute;jar\u0026rsquo; spring, with 38\u0026ordm; C. The hydrochemistry of the low temperature springs shows a predominance of calcium bicarbonate. In contrast, the high temperature springs mainly show silica, chlorides, and sulphates. This correspondence between high water output temperatures and siliceous-sulphate-chloride hydrochemistry appears also in the \u0026gt;\u0026thinsp;30\u0026ordm; C hot springs of the Guadix Basin. The current climatic scenario of the Baza Basin shows an average annual temperature of 13.5\u0026ordm; C (13.8\u0026ordm; C in the Baza station and 13.3\u0026ordm; C in the Orce station; \u003cspan class=\"ExternalRef\"\u003e\u003cspan class=\"RefSource\"\u003ehttps://es.climate-data.org\u003c/span\u003e\u003c/span\u003e), with strong annual oscillations. Specifically, average temperature ranges between 5\u0026ordm; C and 9\u0026ordm; C from November to March, averaging\u0026thinsp;~\u0026thinsp;25\u0026ordm; C during summer. Likewise, the historical record of annual rainfall averages 335 mm at the Baza station and 366 mm at the Orce station (\u003cspan class=\"ExternalRef\"\u003e\u003cspan class=\"RefSource\"\u003ehttps://es.climate-data.org\u003c/span\u003e\u003c/span\u003e). In any case, the average water output temperature of the hot springs in the Baza Basin is 21\u0026ordm; C, which is 7.5\u0026ordm; C higher than the average annual atmospheric temperature in the basin. These data indicate that the water output temperatures of the springs are slightly lower than the average temperatures in summer, but much higher than in autumn and winter. This underlines the thermal character of these hot springs, whose flow rate varies between 1 and 400 l/s.\u003c/p\u003e\n\u003c/div\u003e\n\u003cdiv id=\"Sec4\" class=\"Section2\"\u003e\n \u003ch2\u003e1.3 Tectonic dynamism\u003c/h2\u003e\n \u003cp\u003eFrom a tectonic point of view, the Baza Basin lies in the interior of the alpine-type Betic mountain range, showing three main fracture systems, which are NE-SW, NNW-SSE and NNE-SSW oriented (Sanz de Galdeano, 1983, 1990, \u003cspan class=\"CitationRef\"\u003e2008\u003c/span\u003e; Sanz de Galdeano and Vera, 1991, 1992; Sanz de Galdeano et al., 2007, 2012). These fractures played an important role in the tectono-sedimentary evolution of the Betic mountain range and generated a deep step-like geometry, which is indicative of epicortical extension processes (Sanz de Galdeano and Vera, 1992; Platt et al., \u003cspan class=\"CitationRef\"\u003e2013\u003c/span\u003e). Sanz de Galdeano et al. (2007) established two depocenters in the Baza Basin, induced by NNW-SSE and N70\u0026ordm;E faults that resulted in a regional ENE extension, as deduced by gravity maps of regional, residual and Bouguer anomalies: -150 mGal in Benamaurel (central zone of Baza Basin) and \u0026minus;\u0026thinsp;16 mGal at the north of the Orce village (central zone of Orce sub-basin). Considering basement densities of 2.67 g/cm\u003csup\u003e3\u003c/sup\u003e and 2.32 g/cm\u003csup\u003e3\u003c/sup\u003e for the sedimentary infillings of both sectors, respectively (Sanz de Galdeano et al., 2007), these result in basement depths of ~\u0026thinsp;1300 m and ~\u0026thinsp;500 m, respectively. In this sense, Sanz de Galdeano (\u003cspan class=\"CitationRef\"\u003e2008\u003c/span\u003e) highlighted the Cadiz-Alicante N70\u0026ordm;E fault as the boundary of the inferred depocenters in the central Baza and Orce-North sectors. Likewise, Sanz de Galdeano et al. (2012, 2020) described using geological criteria the main active fractures present in the Baza Basin, which have most likely induced the historical earthquakes recorded in this region: the Galera fault (23 km long and \u0026gt;\u0026thinsp;5 km deep), the Baza fault (37 km long and \u0026gt;\u0026thinsp;10 km deep), the Botardo-Alfahuara fault (22 km long and \u0026gt;\u0026thinsp;5 km deep), the Benamaurel fault (19 km long and \u0026gt;\u0026thinsp;10 km deep), and the C\u0026uacute;llar faults (North and West, ~\u0026thinsp;9 km long and 5\u0026ndash;10 km deep, deduced in this case only by seismic criteria). Seismic phenomena were frequent during the Plio-Pleistocene, as evidenced by the presence of paleo-seismites (Alfaro et al., \u003cspan class=\"CitationRef\"\u003e1997\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2000\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2010\u003c/span\u003e; Gibert et al., \u003cspan class=\"CitationRef\"\u003e2005\u003c/span\u003e). Therefore, considering a standard value of the upper crustal geothermal gradient of 30\u0026ordm; C/km (Harrison et al., \u003cspan class=\"CitationRef\"\u003e1990\u003c/span\u003e; Gupta and Roy, \u003cspan class=\"CitationRef\"\u003e2007\u003c/span\u003e), the depth calculated for these faults and the possible hydrothermal phenomena associated with them, we estimate temperatures of at least 150\u0026ordm;\u0026ndash;300\u0026ordm; C in their deepest zones.\u003c/p\u003e\n\u003c/div\u003e\n\u003cdiv id=\"Sec5\" class=\"Section2\"\u003e\n \u003ch2\u003e1.4 Objectives\u003c/h2\u003e\n \u003cp\u003eThe main goal of this study is to establish a causal link between the presence of hydrothermal upwelling-related deposits in the continental Plio-Pleistocene infillings of the Baza Basin, their tectono-sedimentary origin, and their paleoecological consequences. For doing so, we have: (i) reviewed and documented the tectono-sedimentary context of the Baza Basin; (ii) performed a detailed field study, sampling selected stratigraphic sections; (iii) studied the sedimentology of these sections, analyzing the geochemical and mineralogical composition of the samples; and (iv) performed both local and high scale (Africa and Eurasia) correlations of the relationship between tectonic activity and thermal springs, on the one hand, and the presence of archaeological and paleontological sites, on the other.\u003c/p\u003e\n\u003c/div\u003e"},{"header":"2 Materials and Methods","content":"\u003cp\u003eWe have reviewed and documented the tectono-sedimentary context of the main archaeological and paleontological sites of the Baza Basin, as well as the sedimentary and compositional models of many modern and ancient thermal springs worldwide distributed together with the paleoanthropological sites linked to these springs.\u003c/p\u003e \u003cp\u003eField work in the Baza Basin consisted of geological mapping and observations at a mesoscopic scale of lithologies and their sedimentary structures, as well as of the main faults affecting the sedimentary infillings of the basin. Detailed stratigraphic profiles were obtained and the different sedimentary facies present in them were sampled (Fig.\u0026nbsp;\u003cspan refid=\"Fig2\" class=\"InternalRef\"\u003e2\u003c/span\u003e). Extensive sediment sampling was carried out along the stratigraphic record surveyed (Figs.\u0026nbsp;\u003cspan refid=\"Fig1\" class=\"InternalRef\"\u003e1\u003c/span\u003eB\u0026ndash;C, Fig.\u0026nbsp;\u003cspan refid=\"Fig2\" class=\"InternalRef\"\u003e2\u003c/span\u003e) to perform mineral and chemical analyses for identifying those facies potentially associated with a hydrothermal origin.\u003c/p\u003e \u003cp\u003e \u003c/p\u003e \u003cp\u003eIn this study, we have analyzed 56 samples located throughout the Baza Basin, which correspond to eight types of lithologies: calcilutites, lutites, dark lutites, marls, travertines, cherts, gypsum, and native sulfur. The samples were analyzed geochemically by means of X-ray fluorescence (XRF). In addition, 14 of these samples were selected for analyzing their mineralogy by means of X-ray diffraction (XRD). All measurements were carried out at the Central Services of Support to Research (\u0026lsquo;Servicios Centrales de Apoyo a la Investigaci\u0026oacute;n\u0026rsquo;, SCAI) of the University of M\u0026aacute;laga (UMA).\u003c/p\u003e \u003cdiv id=\"Sec7\" class=\"Section2\"\u003e \u003ch2\u003e2.1 Elemental analysis\u003c/h2\u003e \u003cp\u003eChemical analyses of the abundance of major and some trace elements were carried out using a Thermo Fisher ADVANT'XP wavelength dispersive atomic fluorescence spectrometer (XRF). This equipment consists of an autosampler with capacity for a total of 161 samples, a Rh tube and a goniometer with three collimators and four crystals that allow measuring from F to U. The measurements were obtained in a He atmosphere with the sample in bead. The results were analyzed with Thermofisher's UniQuant software, which allows the semi-quantification of unknown materials without the need of standards or reference materials. Relative errors in the results range between 0.3% and 1.5% in the case of the elements with higher proportions, and up to 5% in the case of those elements with lower proportions.\u003c/p\u003e \u003cp\u003eIn order to evaluate the influence of hydrothermal activity, the ratios (Fe\u0026thinsp;+\u0026thinsp;Mn)/Ti and Al/(Al\u0026thinsp;+\u0026thinsp;Fe\u0026thinsp;+\u0026thinsp;Mn) were calculated (Bostr\u0026ouml;m et al., \u003cspan citationid=\"CR36\" class=\"CitationRef\"\u003e1973\u003c/span\u003e; Murray, \u003cspan citationid=\"CR172\" class=\"CitationRef\"\u003e1994\u003c/span\u003e; Madondo et al., \u003cspan citationid=\"CR148\" class=\"CitationRef\"\u003e2021\u003c/span\u003e; Zhang et al., \u003cspan citationid=\"CR270\" class=\"CitationRef\"\u003e2023\u003c/span\u003e). Moreover, an Al-Fe-Mn ternary diagram was also used to differentiate between hydrothermal, detrital or biogenic origins (Adachi et al., \u003cspan citationid=\"CR2\" class=\"CitationRef\"\u003e1986\u003c/span\u003e).\u003c/p\u003e \u003c/div\u003e \u003cdiv id=\"Sec8\" class=\"Section2\"\u003e \u003ch2\u003e2.2 Mineralogical analysis\u003c/h2\u003e \u003cp\u003eFourteen samples, selected \u003cem\u003ea priori\u003c/em\u003e under a hydrothermal context, were analyzed by X-ray Diffraction (XRD) based on the XRF data. The measurements were obtained with an X'Pert PRO MPD diffractometer from PANalytical and measured in high resolution conditions. The X-ray tube has a Cu anode and the diffractometer in the incident beam consists of a Johansson-type primary monochromator with a Ge (111) crystal, which provides a strictly monochromatic Cu K\u003csub\u003eα1\u003c/sub\u003e radiation. Random powder X-ray diffraction (XRD) of bulk-rocks was used for the identification of mineral phases, measuring the diffraction angles and the intensities of the diffracted beams, which allows to compare the resulting diffraction pattern with a reference database. The comparison was carried out with PDF-4\u0026thinsp;+\u0026thinsp;2021 (Gates-Rector and Blanton, \u003cspan citationid=\"CR95\" class=\"CitationRef\"\u003e2019\u003c/span\u003e). When the acquired data were of sufficient quality (i.e., measured with a wide range of angles, good resolution and high intensity/noise ratio), they were used not only to identify mineral phases but also to quantify them. To determine the proportion of phyllosilicate in the global samples, the phyllosilicates (4.46 \u0026Aring;)/quartz (3.343\u0026Aring;) peak-area ratio was estimated from XRD patterns of randomly oriented powder. D-spacing and the full width at half-high maximum (FWHM) were determined on the (101) reflection peak of the opal specimen in order to establish the type of opal polymorphs (Elzea et al., \u003cspan citationid=\"CR68\" class=\"CitationRef\"\u003e1994\u003c/span\u003e).\u003c/p\u003e \u003c/div\u003e"},{"header":"3 Results","content":"\u003cp\u003eAlmost one hundred archaeological and paleontological sites with vertebrate remains dated between the Early Pliocene (Ruscinian) and the late Early Pleistocene (Galerian) have been described in the Baza Basin (e.g., Casas et al., \u003cspan class=\"CitationRef\"\u003e1975\u003c/span\u003e; Ru\u0026iacute;z-Bustos, 1976; Ru\u0026iacute;z-Bustos, 1991; Cuevas et al., \u003cspan class=\"CitationRef\"\u003e1984\u003c/span\u003e; Agust\u0026iacute;, \u003cspan class=\"CitationRef\"\u003e1985\u003c/span\u003e; Mazo et al., \u003cspan class=\"CitationRef\"\u003e1985\u003c/span\u003e; Soria, \u003cspan class=\"CitationRef\"\u003e1986\u003c/span\u003e; Moy\u0026agrave;-Sol\u0026agrave; et al., 1987; Agust\u0026iacute; et al, \u003cspan class=\"CitationRef\"\u003e1987\u003c/span\u003e; Mart\u0026iacute;n-Su\u0026aacute;rez, \u003cspan class=\"CitationRef\"\u003e1988\u003c/span\u003e; Alberdi and Bonadonna, \u003cspan class=\"CitationRef\"\u003e1989\u003c/span\u003e; Ses\u0026eacute;, \u003cspan class=\"CitationRef\"\u003e1989\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e1991\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2006\u003c/span\u003e; Mart\u0026iacute;nez-Navarro, \u003cspan class=\"CitationRef\"\u003e1991\u003c/span\u003e; Soria and Ruiz-Bustos, \u003cspan class=\"CitationRef\"\u003e1991\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e1992\u003c/span\u003e; Guerra-Merch\u0026aacute;n, \u003cspan class=\"CitationRef\"\u003e1992\u003c/span\u003e; Ses\u0026eacute; \u003cspan class=\"CitationRef\"\u003e1994\u003c/span\u003e; Ses\u0026eacute; and Sevilla, \u003cspan class=\"CitationRef\"\u003e1996\u003c/span\u003e; Gibert and Gibert, \u003cspan class=\"CitationRef\"\u003e2003\u003c/span\u003e; Ros-Montoya, \u003cspan class=\"CitationRef\"\u003e2010\u003c/span\u003e; Maldonado-Garrido et al, \u003cspan class=\"CitationRef\"\u003e2017\u003c/span\u003e; Ros-Montoya et al., \u003cspan class=\"CitationRef\"\u003e2017\u003c/span\u003e, \u003cspan class=\"CitationRef\"\u003e2018\u003c/span\u003e; Mart\u0026iacute;nez-Navarro et al., \u003cspan class=\"CitationRef\"\u003e2018\u003c/span\u003e). Thirty-three of these sites were selected by their archaeological or paleontological relevance, as well as by their clear geographic, stratigraphic and chronological definition (Table \u003cspan class=\"InternalRef\"\u003e1\u003c/span\u003e). Of these sites, 82% (27 out of 33) lie in the Orce sub-basin. Thirteen sites (39%) yielded macrovertebrate remains, with evidence of hominin presence in four of them. These sites were also positioned in a stratigraphic series (Fig. \u003cspan class=\"InternalRef\"\u003e2\u003c/span\u003e). In terms of age, 27% of sites are from the Ruscinian, 6% from the Early Villafranchian, 18% from the Middle Villafranchian, 43% from the Late Villafranchian, 3% from the Epivillafranchian, and the remaining 3% from the Galerian.\u003c/p\u003e\n\u003cp\u003eThe field study showed the presence of 17 levels of cherts, all located in the Orce sub-basin, mainly to the south of the town of Galera (Fig. \u003cspan class=\"InternalRef\"\u003e1\u003c/span\u003eC). Some paleontological sites (e.g., FN-1, GA-2 and NU-2) are situated just a few decimeters above these siliceous deposits (Fig. \u003cspan class=\"InternalRef\"\u003e2\u003c/span\u003e). Chert beds typically occur in irregular shaped strata, with a lateral extent ranging from 7 m to \u0026gt;\u0026thinsp;200 m, and a thickness of 8\u0026ndash;80 cm (Figs. \u003cspan class=\"InternalRef\"\u003e3\u003c/span\u003eA, C). The cartographic relationship between the appearance of cherts and the presence of tectonic fractures is remarkable (Fig. \u003cspan class=\"InternalRef\"\u003e1\u003c/span\u003eC; Figs. \u003cspan class=\"InternalRef\"\u003e3\u003c/span\u003eG, I, J). Lithologically, these cherts appear as white, dark gray or light brown massive facies. At an outcrop scale, they show abundant diaclases, micropores, changes of coloration, and the presence of conchoidal fractures. In some cases, the cherts appear with decimeter to centimeter-scale gypsum mineralizations within them (Fig. \u003cspan class=\"InternalRef\"\u003e3\u003c/span\u003eH). Another remarkable aspect is that these cherts appear in different sedimentary contexts, in stratigraphic contact with other six types of facies: limestones, marls, gypsum, (calci)lutites, and sandstones (Fig. \u003cspan class=\"InternalRef\"\u003e2\u003c/span\u003e).\u003c/p\u003e\n\u003cdiv id=\"Sec10\" class=\"Section2\"\u003e\n \u003ch2\u003e3.1 Elemental analysis\u003c/h2\u003e\n \u003cp\u003eQuantitative analyses of chemical elements were performed (Tables \u003cspan class=\"InternalRef\"\u003e3\u003c/span\u003e and \u003cspan class=\"InternalRef\"\u003e4\u003c/span\u003e). The results were plotted in the Al-Fe-Mn diagram and the two following ratios were calculated: (Fe\u0026thinsp;+\u0026thinsp;Mn)/Ti and Al/(Al\u0026thinsp;+\u0026thinsp;Fe\u0026thinsp;+\u0026thinsp;Mn) (Fig. \u003cspan class=\"InternalRef\"\u003e4\u003c/span\u003e and Table \u003cspan class=\"InternalRef\"\u003e5\u003c/span\u003e). A common feature of hydrothermal deposits is an Al/(Al\u0026thinsp;+\u0026thinsp;Fe\u0026thinsp;+\u0026thinsp;Mn) ratio of \u0026lt;\u0026thinsp;0.4, along with a (Fe\u0026thinsp;+\u0026thinsp;Mn)/Ti ratio of \u0026gt;\u0026thinsp;15 (Bostr\u0026ouml;m et al., \u003cspan class=\"CitationRef\"\u003e1973\u003c/span\u003e; Murray, \u003cspan class=\"CitationRef\"\u003e1994\u003c/span\u003e; Zhang et al., \u003cspan class=\"CitationRef\"\u003e2023\u003c/span\u003e). Forty-two out of the fifty-six samples analyzed did not show mineral and/or geochemical indicators that could \u003cem\u003ea priori\u003c/em\u003e be linked to hydrothermal contexts, while 14 provided preliminary results that are compatible with a hydrothermal origin according to the Al-Fe-Mn diagram and the (Fe\u0026thinsp;+\u0026thinsp;Mn)/Ti - Al/(Al\u0026thinsp;+\u0026thinsp;Fe\u0026thinsp;+\u0026thinsp;Mn) ratios (Bostr\u0026ouml;m et al., \u003cspan class=\"CitationRef\"\u003e1973\u003c/span\u003e; Adachi et al., \u003cspan class=\"CitationRef\"\u003e1986\u003c/span\u003e; Yamamoto, \u003cspan class=\"CitationRef\"\u003e1987\u003c/span\u003e; Murray, \u003cspan class=\"CitationRef\"\u003e1994\u003c/span\u003e; Yamamoto et al., \u003cspan class=\"CitationRef\"\u003e2013\u003c/span\u003e, Fig. \u003cspan class=\"InternalRef\"\u003e4\u003c/span\u003e and Table \u003cspan class=\"InternalRef\"\u003e5\u003c/span\u003e). Eight of these samples (AQ-2, GA-3A, GA-3B, GA-6, GA-7, SX-A, FN1A and CN-1) correspond to cherts, three to travertines (CCH-B, TCV and TE100), two to calcilutites (PL-1 and OR-1) and one to dark lutites (BZ1-A). Except for the dark lutites of the Pliocene site of Baza-1 (Figs. \u003cspan class=\"InternalRef\"\u003e1\u003c/span\u003eB, \u003cspan class=\"InternalRef\"\u003e2\u003c/span\u003e), all the others come from materials in the Orce sub-basin of Late Pliocene to Late Pleistocene age. The (Fe\u0026thinsp;+\u0026thinsp;Mn)/Ti and Al/(Al\u0026thinsp;+\u0026thinsp;Fe\u0026thinsp;+\u0026thinsp;Mn) ratios of these samples are in agreement with conditions of hydrothermalism for the samples of travertines and calcilutites, but not for sample the dark lutites. The last sample provided results of 13.24 and 0.42, respectively. Although these values are very close to the limit of hydrothermal conditions (\u0026gt;\u0026thinsp;15 and \u0026lt;\u0026thinsp;0.4, respectively), they are placed outside the area of hydrothermalism. These geochemical ratios could not be applied to chert samples, because neither Al nor Ti were measured in them.\u003c/p\u003e\n \u003cp\u003eChert samples showed SiO\u003csub\u003e2\u003c/sub\u003e concentrations in excess of 90%, with the exception of two samples, FN1A and GA-6, which exhibited higher concentrations of other compounds (e.g., 14.23% of CaO in FN1A and 5.72% in GA-6). Their variance in composition may be attributed to the presence of calcite in the case of FN1A and gypsum in the case of GA-6, in which a concentration of 6.6% of SO\u003csub\u003e3\u003c/sub\u003e was measured (Table \u003cspan class=\"InternalRef\"\u003e3\u003c/span\u003e). Other trace elements are present in proportions\u0026thinsp;\u0026lt;\u0026thinsp;1% by weight. The geochemical composition of the cherts exhibits an uneven distribution of trace elements (Table \u003cspan class=\"InternalRef\"\u003e4\u003c/span\u003e). Al was absent in all cases, and this was also the case for Mn except for samples CN1 and FN1A, in which this element was recorded with abundances of \u0026lt;\u0026thinsp;0.03%. Fe, Ca, and Cl were consistently detected in all samples. Moreover, Mg, S, K, and Cu were present in every sample except CN1, but the proportions of these elements were lower than in the other samples analyzed in this study.\u003c/p\u003e\n \u003cp\u003eThe five additional samples linked \u003cem\u003ea priori\u003c/em\u003e to a hydrothermal origin include travertines, dark lutites and calcilutites. The travertine levels (samples CCH-B, TCV and TE100, Fig. \u003cspan class=\"InternalRef\"\u003e1\u003c/span\u003eC) are located to the west of the town of Orce. They consist of brown stratiform layers (Fig. \u003cspan class=\"InternalRef\"\u003e3\u003c/span\u003eE) in sedimentary unconformity with the materials deposited below them, which form patchy structures with a lateral development limited to a few meters or, in the case of samples TCV and TE100, to \u0026sim;25 meters. Internal annular carbonate growths developed over previous vegetal structures are visible in these travertines (Fig. \u003cspan class=\"InternalRef\"\u003e3\u003c/span\u003eF). Pisolitic gravels of chaotic aspect and centimeter-scale speleothems are also common. The other travertines present in the Baza Basin show stratiform levels of decimetric thickness in the Venta Micena sector (Fig. \u003cspan class=\"InternalRef\"\u003e2\u003c/span\u003e), where they are stratigraphically close to the paleontological sites of this area, and there are also travertine buildings of metric thickness in the vicinity of currently active hydrothermal springs in the Z\u0026uacute;jar sector (Fig. \u003cspan class=\"InternalRef\"\u003e1\u003c/span\u003eB). According to the dating of similar travertine deposits in the Alic\u0026uacute;n de las Torres current hot spring, which is located in the Guadix Basin (Hern\u0026aacute;ndez et al., \u003cspan class=\"CitationRef\"\u003e2000\u003c/span\u003e; Crespo et al., \u003cspan class=\"CitationRef\"\u003e2014\u003c/span\u003e), the age of these travertines is Middle to Late Pleistocene.\u003c/p\u003e\n \u003cp\u003eThe dark lutites that show a geochemical profile compatible with a hydrothermal origin consist of a\u0026thinsp;~\u0026thinsp;5 cm thick level that is located at 45 cm from the top of the Baza-1 paleontological site (Figs. \u003cspan class=\"InternalRef\"\u003e2\u003c/span\u003e, \u003cspan class=\"InternalRef\"\u003e3\u003c/span\u003eD). From a lithological point of view, these lutites show dark gray tones and contain sub-millimeter-scale parallel laminations, root remains, small fragments of gastropod shells, oxidations, and micas randomly distributed in the sediment. The presence of dark lutites in the Baza Basin is commonly recorded in thin layers present in most of the stratigraphic series of Plio-Pleistocene age (Fig. \u003cspan class=\"InternalRef\"\u003e2\u003c/span\u003e). Mesoscopic scale sedimentary structures of traction origin and linked to detrital deposits (e.g., sandy microlevels, coarsening/finning upward sequences, imbrications, or scour-marks) are absent. Analyses of sample BZ1-A (Tables \u003cspan class=\"InternalRef\"\u003e3\u003c/span\u003e\u0026ndash;\u003cspan class=\"InternalRef\"\u003e4\u003c/span\u003e) showed a main chemical composition of Si\u0026thinsp;\u0026gt;\u0026thinsp;Fe\u0026thinsp;\u0026gt;\u0026thinsp;Al\u0026thinsp;\u0026gt;\u0026thinsp;Ca\u0026thinsp;\u0026gt;\u0026thinsp;K\u0026thinsp;\u0026gt;\u0026thinsp;Mg. Compared to other dark lutites studied, BZ1-A stands out for its notably higher concentration of Fe\u003csub\u003e2\u003c/sub\u003eO\u003csub\u003e3\u003c/sub\u003e, which reaches 11.19%. This iron content exceeds those of other samples of dark lutites. Additionally, this sample exhibits elevated levels of K, Mn, S, Cl, Ba, Cu, Rb, Pb, Sn, and Cs compared to the average contents of the samples of dark lutites.\u003c/p\u003e\n \u003cp\u003eCalcilutites are present in all stratigraphic profiles except Z\u0026uacute;jar (Fig. \u003cspan class=\"InternalRef\"\u003e2\u003c/span\u003e) and use to appear as light-colored levels of decimetric to metric thickness. From a lithological point of view, they typically exhibit massive facies or scarce internal arrangement, although sometimes there are internal shallowing-upward sequences with edaphogenic-type facies of reddish tones situated at their top. The PL-1 calcilutite sample shows a main geochemical composition of Ca\u0026thinsp;\u0026gt;\u0026thinsp;Si\u0026thinsp;\u0026gt;\u0026thinsp;Fe\u0026thinsp;\u0026gt;\u0026thinsp;Al\u0026thinsp;\u0026gt;\u0026thinsp;Mg (Tables \u003cspan class=\"InternalRef\"\u003e3\u003c/span\u003e\u0026ndash;\u003cspan class=\"InternalRef\"\u003e4\u003c/span\u003e). The rest of the elements detected appeared with concentrations\u0026thinsp;\u0026lt;\u0026thinsp;0.6%. In comparison with other calcilutites of the basin, this sample has the highest CaO values and the lowest of SiO\u003csub\u003e2\u003c/sub\u003e, Al\u003csub\u003e2\u003c/sub\u003eO\u003csub\u003e3\u003c/sub\u003e, K\u003csub\u003e2\u003c/sub\u003eO, TiO\u003csub\u003e2\u003c/sub\u003e, Na\u003csub\u003e2\u003c/sub\u003eO, and SO\u003csub\u003e3\u003c/sub\u003e. Finally, the OR-1 calcilutite sample shows a main geochemical composition of Si\u0026thinsp;\u0026gt;\u0026thinsp;Ca\u0026thinsp;\u0026gt;\u0026thinsp;Mn\u0026thinsp;\u0026gt;\u0026thinsp;Al\u0026thinsp;\u0026gt;\u0026thinsp;Fe\u0026thinsp;\u0026gt;\u0026thinsp;Mg. The rest of the elements detected appear with concentrations\u0026thinsp;\u0026lt;\u0026thinsp;0.7%. This sample exhibits the highest MnO value (~\u0026thinsp;10 times more) compared to the average of the calcilutite samples. Additionally, geochemical analyses revealed the presence of some trace elements in the samples (Table \u003cspan class=\"InternalRef\"\u003e4\u003c/span\u003e). Of these, Ba stands out, being exclusively detected in four samples \u003cem\u003ea priori\u003c/em\u003e linked to hydrothermal environments: BZ1-A (dark lutites), OR-1 (calcilutites), TCV (travertines), and TE100 (travertines).\u003c/p\u003e\n\u003c/div\u003e\n\u003cdiv id=\"Sec11\" class=\"Section2\"\u003e\n \u003ch2\u003e3.2 Mineralogical analysis\u003c/h2\u003e\n \u003cp\u003eThe XRD data of cherts show that opal is the principal phase (Table \u003cspan class=\"InternalRef\"\u003e6\u003c/span\u003e, Fig. \u003cspan class=\"InternalRef\"\u003e5\u003c/span\u003e). The d\u003csub\u003e(101)\u003c/sub\u003e (\u0026gt;\u0026thinsp;4.07 \u0026Aring;) and the full width at half-high maximum (FWHM) reflection peak intensity of opal (101) indicate that CT is the polymorph of the opal (Elzea et al., \u003cspan class=\"CitationRef\"\u003e1994\u003c/span\u003e). Most of the samples taken in Galera showed the presence of gypsum in the XRD data, as evidenced by geochemistry (high CaO and SO\u003csub\u003e3\u003c/sub\u003e contents, Table \u003cspan class=\"InternalRef\"\u003e3\u003c/span\u003e) and field observations (Fig. \u003cspan class=\"InternalRef\"\u003e3\u003c/span\u003eH). The presence of silicates and carbonates in some samples (e.g., CN-1, GA-3B and SX-A) may come from the remains of the sedimentary materials above and/or below the levels where these samples were obtained, due to the irregular morphology of the chert strata. Travertine samples mainly consist of calcite (Table \u003cspan class=\"InternalRef\"\u003e6\u003c/span\u003e) and, to a lesser extent, pyrolusite. In addition, other mineral species such as gypsum and halite were found. The XRD data from these samples agree with the expectations from geochemical (Tables \u003cspan class=\"InternalRef\"\u003e3\u003c/span\u003e\u0026ndash;\u003cspan class=\"InternalRef\"\u003e4\u003c/span\u003e) and lithological data. The analyses carried out on the calcilutites of samples PL-1 and OR-1 (Table \u003cspan class=\"InternalRef\"\u003e6\u003c/span\u003e) showed a mineral composition of calcite, quartz and phyllosilicates (muscovite and paragonite). In addition, dolomite and plagioclase were both present as trace amounts. Finally, the geochemistry of the sample of dark lutites with hydrothermal markers (BZ1-A) consisted mainly of phyllosilicates (\u0026gt;\u0026thinsp;50%: muscovite, paragonite, kaolinite, and chlorite) and quartz (Table \u003cspan class=\"InternalRef\"\u003e6\u003c/span\u003e). Accessory minerals included feldspars, carbonates, lepidocrocite, halite, and gypsum. The high FeO\u003csub\u003e3\u003c/sub\u003e, CaO, SO\u003csub\u003e3\u003c/sub\u003e, and Cl (Tables \u003cspan class=\"InternalRef\"\u003e3\u003c/span\u003e and \u003cspan class=\"InternalRef\"\u003e4\u003c/span\u003e) contents justified the presence of lepidocrocite, gypsum and halite.\u003c/p\u003e\n\u003c/div\u003e"},{"header":"4 Discussion","content":"\u003cp\u003eOn a cartographic scale, 14 samples linked to hydrothermal environments were found in the vicinity of traces of the main tectonic fractures present in the basin (Sanz de Galdeano et al., 2007, 2012; Garc\u0026iacute;a-Tortosa et al., \u003cspan citationid=\"CR94\" class=\"CitationRef\"\u003e2011\u003c/span\u003e; Figs.\u0026nbsp;\u003cspan refid=\"Fig1\" class=\"InternalRef\"\u003e1\u003c/span\u003eB\u0026ndash;C): samples AQ-2, GA-3A, GA-3B, GA-6, GA-7, and PL-1 are close to the NE-SW Galera Fault; sample CN1 is close to the NW-SE Alfahuara-Botardo Fault; sample BZ-1A is close to the NNW-SSE Baza Fault; and samples SX-A, FN1A, OR-1, CCH-B, TCV, and TE100 are all in the vicinity of WNW-ESE oriented fractures. This suggests the connection between materials with evidence of hydrothermal activity and their position close to tectonic fractures, as evidenced by the fact that the current thermal springs in the basin are found on or in the vicinity of traces of tectonic fractures (Fig.\u0026nbsp;\u003cspan refid=\"Fig1\" class=\"InternalRef\"\u003e1\u003c/span\u003eB). At an outcrop scale, the relationship between chert deposits and tectonic fractures is evident (Figs.\u0026nbsp;\u003cspan refid=\"Fig3\" class=\"InternalRef\"\u003e3\u003c/span\u003eG, I, J).\u003c/p\u003e \u003cdiv id=\"Sec13\" class=\"Section2\"\u003e \u003ch2\u003e4.1 Origin of cherts and other facies potentially generated in hydrothermal contexts\u003c/h2\u003e \u003cp\u003eChert layers appear in the NE sector of the Baza Basin (Orce sub-basin) and are cartographically associated with fracture traces. In fact, a chert dyke (Fig.\u0026nbsp;\u003cspan refid=\"Fig3\" class=\"InternalRef\"\u003e3\u003c/span\u003eI) was recognized to the SE of the town of Galera, coincident with the trace of a NE-SW fault that shows synsedimentary features (S\u0026aacute;nchez-Roa et al., \u003cspan citationid=\"CR217\" class=\"CitationRef\"\u003e2016\u003c/span\u003e). In the case of Fuente Nueva-1 and Galera, the chert deposits are found near levels rich in paleontological remains (Fig.\u0026nbsp;\u003cspan refid=\"Fig2\" class=\"InternalRef\"\u003e2\u003c/span\u003e).\u003c/p\u003e \u003cp\u003eHydrothermal deposits often show high concentrations of iron and manganese, while aluminum and titanium are reliable indicators of sedimentary input (Bostr\u0026ouml;m et al., \u003cspan citationid=\"CR36\" class=\"CitationRef\"\u003e1973\u003c/span\u003e; Adachi et al., \u003cspan citationid=\"CR2\" class=\"CitationRef\"\u003e1986\u003c/span\u003e; Murray, \u003cspan citationid=\"CR172\" class=\"CitationRef\"\u003e1994\u003c/span\u003e; Zhang et al., \u003cspan citationid=\"CR270\" class=\"CitationRef\"\u003e2023\u003c/span\u003e). This is why many research studies use the ratios (Fe\u0026thinsp;+\u0026thinsp;Mn)/Ti and Al/(Al\u0026thinsp;+\u0026thinsp;Fe\u0026thinsp;+\u0026thinsp;Mn) to identify hydrothermal influence (Table\u0026nbsp;\u003cspan refid=\"Tab5\" class=\"InternalRef\"\u003e5\u003c/span\u003e). The application of the Al-Fe-Mn diagram (Adachi et al., \u003cspan citationid=\"CR2\" class=\"CitationRef\"\u003e1986\u003c/span\u003e; Yamamoto, \u003cspan citationid=\"CR264\" class=\"CitationRef\"\u003e1987\u003c/span\u003e; Yamamoto et al., \u003cspan citationid=\"CR265\" class=\"CitationRef\"\u003e2013\u003c/span\u003e) to the chert samples (Fig.\u0026nbsp;\u003cspan refid=\"Fig4\" class=\"InternalRef\"\u003e4\u003c/span\u003e) showed a geochemical composition in agreement with a hydrothermal origin. Although Al-Fe-Mn diagrams have been applied to deposits of marine origin (Adachi et al., \u003cspan citationid=\"CR2\" class=\"CitationRef\"\u003e1986\u003c/span\u003e; Yamamoto, \u003cspan citationid=\"CR264\" class=\"CitationRef\"\u003e1987\u003c/span\u003e; Boncheva et al., \u003cspan citationid=\"CR34\" class=\"CitationRef\"\u003e2023\u003c/span\u003e; El-Moghazy et al., \u003cspan citationid=\"CR67\" class=\"CitationRef\"\u003e2023\u003c/span\u003e; Jurkowska and Swierczewska-Gladysz, 2024; among others), they have also been applied to continental environments (Madondo et al., \u003cspan citationid=\"CR148\" class=\"CitationRef\"\u003e2021\u003c/span\u003e; Zhou et al., \u003cspan citationid=\"CR271\" class=\"CitationRef\"\u003e2022\u003c/span\u003e). The results obtained in the analysis of the chert samples from the Baza Basin are compatible with a hydrothermal origin for them if we consider other evidence (see below) and also according to the Al-Fe-Mn contents and main trace elements present in the siliceous rocks of the North Pacific (Adachi et al., \u003cspan citationid=\"CR2\" class=\"CitationRef\"\u003e1986\u003c/span\u003e). On the opposite, the (Fe\u0026thinsp;+\u0026thinsp;Mn)/Ti and Al/(Al\u0026thinsp;+\u0026thinsp;Fe\u0026thinsp;+\u0026thinsp;Mn) ratios have been used in sediments to determine their hydrothermal origin (Bostr\u0026ouml;m et al., \u003cspan citationid=\"CR36\" class=\"CitationRef\"\u003e1973\u003c/span\u003e; Murray, \u003cspan citationid=\"CR172\" class=\"CitationRef\"\u003e1994\u003c/span\u003e; Zhang et al., \u003cspan citationid=\"CR270\" class=\"CitationRef\"\u003e2023\u003c/span\u003e). Therefore, the data obtained from the chert samples are expected to be more reliable than those derived from samples without chert contents. In any case, the use of both methods is complementary and the comparison of their results in these continental environments allows their evaluation as proxies in this sedimentary context. The main mineral composition of opal-CT in the chert samples indicates a silica mineral phase linked to hydrothermal conditions (Kano and Taguchi, \u003cspan citationid=\"CR130\" class=\"CitationRef\"\u003e1982\u003c/span\u003e; Fl\u0026ouml;rke et al., \u003cspan citationid=\"CR73\" class=\"CitationRef\"\u003e1990\u003c/span\u003e; Lynne et al., \u003cspan citationid=\"CR147\" class=\"CitationRef\"\u003e2005\u003c/span\u003e; Fr\u0026ouml;hlich, \u003cspan citationid=\"CR79\" class=\"CitationRef\"\u003e2020\u003c/span\u003e).\u003c/p\u003e \u003cp\u003eSilica-rich deposits linked to \u0026gt;\u0026thinsp;250\u0026ordm;C hydrothermal fluids have been associated with tectonic fractures in Permian cherts based on their geochemical analyses and Al-Fe-Mn diagrams (for review and references, see Zhou et al., \u003cspan citationid=\"CR271\" class=\"CitationRef\"\u003e2022\u003c/span\u003e). According to the Na/K calculation method (Fournier and Truesdell, \u003cspan citationid=\"CR78\" class=\"CitationRef\"\u003e1973\u003c/span\u003e; Fournier, \u003cspan citationid=\"CR76\" class=\"CitationRef\"\u003e1977\u003c/span\u003e; Giggenbach et al., \u003cspan citationid=\"CR103\" class=\"CitationRef\"\u003e1988\u003c/span\u003e), the estimates obtained with geothermometers for the high-temperature hot springs that are currently active in the Guadix-Baza Depression are 88\u0026ndash;89\u0026ordm;C. A similar situation occurs in Alhama de Granada, SE Spain (L\u0026oacute;pez-Chicano et al., \u003cspan citationid=\"CR144\" class=\"CitationRef\"\u003e2001\u003c/span\u003e). Moreover, the hot springs of the Guadix-Baza Depression (Diputaci\u0026oacute;n de Granada-ITGE, \u003cspan citationid=\"CR64\" class=\"CitationRef\"\u003e1990\u003c/span\u003e; Table\u0026nbsp;\u003cspan refid=\"Tab2\" class=\"InternalRef\"\u003e2\u003c/span\u003e) show a direct relationship between their output water temperature and silica contents (Garc\u0026iacute;a-Aguilar et al., \u003cspan citationid=\"CR90\" class=\"CitationRef\"\u003e2014\u003c/span\u003e), which can reach up to 54 mg/l at 44\u0026ordm; C in the Graena spring. According to the standard values of the upper crustal geothermal gradient, tectonic and geophysical gravimetry data, Sanz de Galdeano et al. (2007) suggested a temperature range in the deep traces of the main faults of the Baza Basin of 150\u0026ndash;300\u0026ordm;C, which agrees with a hydrothermal scenario. These temperatures are compatible with those inferred for the formation of Permian chert nodules in the Junggar Basin (Zhou et al., \u003cspan citationid=\"CR271\" class=\"CitationRef\"\u003e2022\u003c/span\u003e).\u003c/p\u003e \u003cp\u003eThe trace-element contents of the cherts of the Baza Basin are very variable depending on the sample considered, because each of them shows a particular geochemical profile. The case of sample FN1A, which is located a few decimeters below the Fuente Nueva-1 site, stands out for its variety of trace elements, many of them not recorded in other deposits (e.g., Sr, As, Mo, and W; Table\u0026nbsp;\u003cspan refid=\"Tab4\" class=\"InternalRef\"\u003e4\u003c/span\u003e). In this sense, the presence of As has been associated with hydrothermal fluids near tectonic fractures west of the Orce sub-basin (S\u0026aacute;nchez-Roa et al., \u003cspan citationid=\"CR217\" class=\"CitationRef\"\u003e2016\u003c/span\u003e) and the presence of Sr has also been linked to hydrothermal contexts in the Plio-Pleistocene lacustrine environments of the Guadix-Baza Depression (Garc\u0026iacute;a-Aguilar et al., \u003cspan citationid=\"CR90\" class=\"CitationRef\"\u003e2014\u003c/span\u003e). On the other hand, the variety of geochemical profiles found in the chert samples is compatible with the variety of hydrochemical profiles (both qualitatively and quantitatively) and output temperatures present in the hot springs that are currently active in the basin (Table\u0026nbsp;\u003cspan refid=\"Tab2\" class=\"InternalRef\"\u003e2\u003c/span\u003e). These hydrochemical profiles must be linked to the geometry of the tectonic fractures associated with each hot spring and to the different lithological substrates that the hydrothermal fluids passed through on their way to the surface. This cause-effect relationship could be extrapolated to the ancient Plio-Pleistocene hydrothermal upwellings in the Baza Basin, whose salt contents, output temperatures and water flow would have varied depending on the tectonic, lithological, and structural features present in each case.\u003c/p\u003e \u003cp\u003eThe hydrochemical composition of the hot springs that are currently active in the basin with output temperatures of \u0026gt;\u0026thinsp;35\u0026ordm;C (Cruz-Sanjuli\u0026aacute;n et al., \u003cspan citationid=\"CR55\" class=\"CitationRef\"\u003e1972\u003c/span\u003e; Cruz-Sanjuli\u0026aacute;n and Garc\u0026iacute;a-Rosell, \u003cspan citationid=\"CR54\" class=\"CitationRef\"\u003e1972\u003c/span\u003e, \u003cspan citationid=\"CR56\" class=\"CitationRef\"\u003e1975\u003c/span\u003e; Diputaci\u0026oacute;n de Granada-ITGE, \u003cspan citationid=\"CR64\" class=\"CitationRef\"\u003e1990\u003c/span\u003e; Table\u0026nbsp;\u003cspan refid=\"Tab2\" class=\"InternalRef\"\u003e2\u003c/span\u003e) shows silica contents of \u0026gt;\u0026thinsp;40 mg/l, high sulfate values and the presence of chlorides. Similar conditions are widely documented in hydrothermally active regions, such as the Dead Sea-Jordan River Basin in Israel (Levitte and Eckstein, \u003cspan citationid=\"CR140\" class=\"CitationRef\"\u003e1978\u003c/span\u003e), Yellowstone in USA (Nordstrom et al., \u003cspan citationid=\"CR173\" class=\"CitationRef\"\u003e2005\u003c/span\u003e), Mount Tafta in Iran (Shakeri et al., \u003cspan citationid=\"CR236\" class=\"CitationRef\"\u003e2008\u003c/span\u003e) or Denizli in Turkey (\u0026Ouml;zler, \u003cspan citationid=\"CR181\" class=\"CitationRef\"\u003e2000\u003c/span\u003e), among others. On the other hand, White et al. (\u003cspan citationid=\"CR259\" class=\"CitationRef\"\u003e1956\u003c/span\u003e) and Fournier and Rowe (\u003cspan citationid=\"CR77\" class=\"CitationRef\"\u003e1966\u003c/span\u003e) established the hydrochemical conditions linked to the solubility and subsequent precipitation of silica in thermal waters. They defined a solubility of 315 mg/l for silica at 90\u0026ordm;C and 110 mg/l at 25\u0026ordm; C, demonstrating a correspondence between the output temperature of the hot springs and their silica contents. The rapid rise of hot waters to the surface results in silica oversaturation due to cooling and degasification. Dissolution by high-temperature waters of deep rocks and their subsequent ascent through tectonic fractures is common. In the case of cherts, however, these hot springs should have shown much higher output temperatures (\u0026gt;\u0026thinsp;35\u0026ordm;C) than in the atmosphere. Considering the climatic conditions inferred for the Early Pleistocene in the Baza Basin, these thermal pools would have originated small areas with a stable and warm microclimate throughout the year, suitable for the ecological development of a diverse mammalian community, especially during the cold periods of the Pleistocene (Lisiecki and Raymo, \u003cspan citationid=\"CR141\" class=\"CitationRef\"\u003e2007\u003c/span\u003e; Paillard, \u003cspan citationid=\"CR182\" class=\"CitationRef\"\u003e2017\u003c/span\u003e).\u003c/p\u003e \u003cp\u003eUnder these conditions, cherts originated from the dissolution at depth conditions of rocks with elevated siliceous contents (e.g., quartzites and mica-schists of the Internal Zones of the Betic mountain range; Baena P\u0026eacute;rez et al., \u003cspan citationid=\"CR21\" class=\"CitationRef\"\u003e1979\u003c/span\u003e; Mart\u0026iacute;n Garc\u0026iacute;a et al., \u003cspan citationid=\"CR152\" class=\"CitationRef\"\u003e1980\u003c/span\u003e; Villalobos et al., \u003cspan citationid=\"CR256\" class=\"CitationRef\"\u003e2006\u003c/span\u003e) in high-temperature fluids. According to the most plausible calculated Na/K geothermometers (88\u0026ndash;89\u0026ordm;C) and considering the standard geothermal gradient in the upper crust (25\u0026ndash;30\u0026ordm;C/km, Harrison et al., \u003cspan citationid=\"CR115\" class=\"CitationRef\"\u003e1990\u003c/span\u003e; Gupta and Roy, \u003cspan citationid=\"CR111\" class=\"CitationRef\"\u003e2007\u003c/span\u003e), the estimated depth in the Baza Basin for the dissolution of silica agrees with the basement depth calculated from seismic profiles, 2150 m (Alfaro et al., \u003cspan citationid=\"CR11\" class=\"CitationRef\"\u003e2008\u003c/span\u003e). This depth is similar to the estimation for present-day hot springs in Algeria (Belhai et al., \u003cspan citationid=\"CR28\" class=\"CitationRef\"\u003e2017\u003c/span\u003e) and indicates the presence of a deep fracture network affecting the basin substrate in the Guadix-Baza Depression, where the hot springs would dissolve components of the rocks bound to them. These depth values linked to the origin of thermal upwellings in the basin are lower than the depth range of the tectonic fractures in the basin (5\u0026ndash;10 km), which has been estimated using geological and geophysical criteria (Sanz de Galdeano et al. 2012, 2020).\u003c/p\u003e \u003cp\u003eTravertine levels of decimetric thickness and associated with a hydrothermal context have been detected in the southern bank of the Orce River, west to the town of Orce. The application of the Al-Fe-Mn diagram and the (Fe\u0026thinsp;+\u0026thinsp;Mn)/Ti and Al/(Al\u0026thinsp;+\u0026thinsp;Fe\u0026thinsp;+\u0026thinsp;Mn) ratios to these samples showed a composition compatible with a hydrothermal origin (Fig.\u0026nbsp;\u003cspan refid=\"Fig4\" class=\"InternalRef\"\u003e4\u003c/span\u003e, Table\u0026nbsp;\u003cspan refid=\"Tab5\" class=\"InternalRef\"\u003e5\u003c/span\u003e).\u003c/p\u003e \u003cp\u003eFurthermore, the Al-Fe-Mn diagram suggests that sample BZ1-A of dark lutites may have had some hydrothermal influence. Moreover, the mineralogy of this sample (Table\u0026nbsp;\u003cspan refid=\"Tab6\" class=\"InternalRef\"\u003e6\u003c/span\u003e) indicates a primarily detrital deposit (quartz, paragonite and chlorite) with halite precipitation due to evaporation processes. It is noteworthy that the presence of Sr and Mo was detected in this sample (Table\u0026nbsp;\u003cspan refid=\"Tab4\" class=\"InternalRef\"\u003e4\u003c/span\u003e), as these elements have been found under a hydrothermal context in the Galera deposits (S\u0026aacute;nchez-Roa et al., \u003cspan citationid=\"CR217\" class=\"CitationRef\"\u003e2016\u003c/span\u003e). Despite having very high iron contents, the hydrothermal ratios (Fe\u0026thinsp;+\u0026thinsp;Mn)/Ti and Al/(Al\u0026thinsp;+\u0026thinsp;Fe\u0026thinsp;+\u0026thinsp;Mn)) suggest that this level has not a hydrothermal origin. Given the discrepancy between the Al-Fe-Mn diagram and these ratios, it is important to note that the diagram was designed for analyzing siliceous materials (Adachi et al., \u003cspan citationid=\"CR2\" class=\"CitationRef\"\u003e1986\u003c/span\u003e) while the ratios are typically used for sediments (You et al., \u003cspan citationid=\"CR267\" class=\"CitationRef\"\u003e2020\u003c/span\u003e; Zhang et al., \u003cspan citationid=\"CR270\" class=\"CitationRef\"\u003e2023\u003c/span\u003e). As a result, it is plausible that the ratios provide a more accurate indication of hydrothermalism in these dark lutites. However, although the values in these ratios are close to those expected in hydrothermal environments, they are not within the hydrothermal zone. This casts doubts on whether BZ1-A was influenced by hydrothermalism. In any case, it is remarkable that BZ1-A is the only sample among the 17 samples of dark lutites analyzed that fits the diagram of hydrothermalism.\u003c/p\u003e \u003cp\u003eThe calcilutite samples that provided geochemical indicators of hydrothermal activity correspond to two metric scale levels located near the localities of Hu\u0026eacute;scar (sample PL-1) and Orce (sample OR-1, Fig.\u0026nbsp;\u003cspan refid=\"Fig2\" class=\"InternalRef\"\u003e2\u003c/span\u003e). In both cases, they are placed in the vicinity of paleontological sites with macrofauna. The application of the (Fe\u0026thinsp;+\u0026thinsp;Mn)/Ti and Al/(Al\u0026thinsp;+\u0026thinsp;Fe\u0026thinsp;+\u0026thinsp;Mn) ratios to these samples (Table\u0026nbsp;\u003cspan refid=\"Tab5\" class=\"InternalRef\"\u003e5\u003c/span\u003e) shows values that are compatible with a hydrothermal origin for them. In the case of sample PL-1, the Al-Fe-Mn diagram (Fig.\u0026nbsp;\u003cspan refid=\"Fig4\" class=\"InternalRef\"\u003e4\u003c/span\u003e) shows further evidence of its hydrothermal origin. From the mineralogical point of view, the composition of these samples shows quartz, calcite, dolomite, muscovite, paragonite, and plagioclase (Table\u0026nbsp;\u003cspan refid=\"Tab6\" class=\"InternalRef\"\u003e6\u003c/span\u003e). This association of minerals can be considered detrital-inherited by sedimentary processes, as evidenced by the lithological characteristics observed in the outcrops where the samples were taken. In addition to the previous arguments, the presence of significant amounts of Ba in samples BZ1-A (dark lutites), OR-1 (calcilutites), TCV (travertines), and TE100 (travertines) adds further evidence on the hydrothermal context associated with their deposition (Jankovic, 1982; Canet et al., \u003cspan citationid=\"CR45\" class=\"CitationRef\"\u003e2005\u003c/span\u003e).\u003c/p\u003e \u003cp\u003eThe hydrothermal markers in samples Pl-1, OR-1 and BZ1-A, whose lithologies contain inherited detrital elements, could be interpreted as due to minerals originating from source areas with metamorphic substrates. However, these hydrothermal markers do not appear in other deposits that are lithologically and sedimentologically similar to these three samples, which indicates that the high Fe and Mn contents should have a hydrothermal origin. Concerning the sedimentary context of the cherts and other deposits linked to hydrothermal conditions and their association with archaeological and paleontological sites, it is worth noting that these levels frequently appear on the top and/or bottom of the sites as well as in lateral connection with facies linked to a diversity of sedimentary scenarios.\u003c/p\u003e \u003c/div\u003e \u003cdiv id=\"Sec14\" class=\"Section2\"\u003e \u003ch2\u003e4.2 Paleogeographical reconstruction of hydrothermal activity in the basin\u003c/h2\u003e \u003cp\u003eThe spatio-temporal correlation between the coordinates of the 14 samples with evidence of past hydrothermal activity and the position of the hot springs that are currently active in the Baza Basin allows us to propose a general model of paleogeographic evolution of the basin in relation to its hydrothermal activity (Fig.\u0026nbsp;\u003cspan refid=\"Fig6\" class=\"InternalRef\"\u003e6\u003c/span\u003e). This model makes it possible to specify the cause-effect relationships between the tectonic dynamism of the basin and the occurrence of hydrothermal phenomena linked to tectonic fractures (Figs.\u0026nbsp;\u003cspan refid=\"Fig1\" class=\"InternalRef\"\u003e1\u003c/span\u003eB\u0026ndash;C) at specific places and times that generated in each case different geochemical and mineral markers in the sedimentary environments. These environments can be very diverse, since they do not show a direct genetic relationship with the position of the hydrothermal upwellings. In this way, the cherts, travertines, calcilutites, and dark lutites (i.e., the facies in which hydrothermal markers have been detected) correspond to different sedimentary environments, which include lacustrine zones with some depth (calcilutites), marshy environments (dark lutites), occasional upwellings in favor of tectonic fractures (cherts) and waterfalls or pools of thermal waters (travertines), according with the most plausible sedimentary models (Reading, \u003cspan citationid=\"CR201\" class=\"CitationRef\"\u003e1978\u003c/span\u003e; Prothero and Schwab, \u003cspan citationid=\"CR200\" class=\"CitationRef\"\u003e2004\u003c/span\u003e). Among these sedimentary environments, those linked to the cherts could be considered the only ones with a direct causal relationship between these facies and the presence of hydrothermal activity.\u003c/p\u003e \u003cp\u003e \u003c/p\u003e \u003cp\u003eAs shown in Fig.\u0026nbsp;\u003cspan refid=\"Fig6\" class=\"InternalRef\"\u003e6\u003c/span\u003e, hydrothermal phenomena appear discontinuously in the Baza Basin from the Late Pliocene (Ruscinian) to the present. In this sense, there seems to be a gap between the Upper Ruscinian and the Middle Villafranchian, together with a gap in the lower Late Villafranchian. However, the presence of hydrothermal activity during these periods cannot be ruled out. Its demonstration would depend on a wide sampling of the different sedimentary facies linked to these periods, preferably located near tectonic fractures. Regarding the geographical distribution, it is evident that hydrothermal activity concentrates in the Galera-Orce-Ca\u0026ntilde;ada del Negro sector (Orce sub-basin). This points to an intense tectonic activity in this sub-basin during the Plio-Pleistocene, particularly in four intervals: Late Ruscinian, Middle Villafranchian, upper Late Villafranchian and Galerian to present, with the presence of hydrothermal phenomena that induced mineral and/or geochemical markers in the sedimentary environments associated to these places and times. Likewise, the Baza area seems to show a similar pattern to the Orce sub-basin for the Ruscinian, associated with an important tectonic fracture, the Baza Fault. It is also evident that each node of hydrothermal activity seems to demonstrate a particular physico-chemical characteristic, as shown by the compositional variety of the deposits linked to this activity. In this regard, factors such as the hydrochemical composition and physical parameters of the hydrothermal fluids, the sedimentary environments where they acted and (micro)biological factors would give rise to a wide range of situations. This pattern could have played an important role in the development of local ecosystems in the vicinity of the nodes of hydrothermal activity. Finally, it is evident how these hydrothermal phenomena coincide quite closely in their spatio-temporal coordinates with the position of important paleontological sites, as happens in the case of BZ-1, FN1, HU-1, and BO-7 (Fig.\u0026nbsp;\u003cspan refid=\"Fig2\" class=\"InternalRef\"\u003e2\u003c/span\u003e). If we further extend these spatio-temporal coordinates to a wider range, we see other important sites (e.g., BL, FN-3 and VM) that appear in a geographic and/or chronostratigraphic position close to the nodes of hydrothermal activity. This allows us to establish a possible paleoecological relationship between the presence of hydrothermal phenomena and the occurrence of paleontological sites in the Baza Basin.\u003c/p\u003e \u003c/div\u003e \u003cdiv id=\"Sec15\" class=\"Section2\"\u003e \u003ch2\u003e4.3 Paleoecological implications of hydrothermal activity in the basin\u003c/h2\u003e \u003cp\u003eA contribution of thermal waters would play a role in maintaining during the dry season a minimum level in the water table of the lacustrine environments of the Baza Basin. This would in turn allow the development of a luxurious vegetational cover in the surroundings of the hot springs, which would sustain a large vertebrate community. The hot springs of the Baza Basin led to the appearance of thermal pools of variable extent and length, with waters that differed from the main lacustrine systems in their composition and physical properties. These hot springs would support in part the ecological pyramid of the Baza Basin in a specific network of spatio-temporal nodes, concentrating the plants and animals in their surroundings, which would in turn result in a concentration of ungulate carcasses in the vicinity of the hot springs (Fig.\u0026nbsp;\u003cspan refid=\"Fig7\" class=\"InternalRef\"\u003e7\u003c/span\u003e). This is reflected in some of the most important deposits of the basin, such as the Pliocene sites of Baza-1 and Fuente Nueva-1, which are both linked to a level of dark lutites and to a level of cherts, respectively. From a sedimentary point of view, this provides evidence on the role of hydrothermal events as hot spots for the concentration of terrestrial fauna.\u003c/p\u003e \u003cp\u003e \u003c/p\u003e \u003cp\u003eThe relationship between some ecological factors linked to contexts with thermal waters is shown in Fig.\u0026nbsp;\u003cspan refid=\"Fig8\" class=\"InternalRef\"\u003e8\u003c/span\u003eA, which depicts a direct, statistically significant relationship between prey biomass (kg/km\u003csup\u003e2\u003c/sup\u003e) and annual rainfall (mm) values in a set of African Natural Parks and Game Reserves [data compiled from Bell (\u003cspan citationid=\"CR29\" class=\"CitationRef\"\u003e1981\u003c/span\u003e) and Hatton et al. (\u003cspan citationid=\"CR116\" class=\"CitationRef\"\u003e2015\u003c/span\u003e); see also Coe et al. (\u003cspan citationid=\"CR49\" class=\"CitationRef\"\u003e1976\u003c/span\u003e) for a similar relationship]. However, there is considerable scatter around the least-squares regression line, which primarily depends on soil type and, secondarily, on the presence of thermal springs (Fig.\u0026nbsp;\u003cspan refid=\"Fig8\" class=\"InternalRef\"\u003e8\u003c/span\u003eB), which increase the net primary productivity thanks to their saline contents (Garc\u0026iacute;a-Aguilar et al., \u003cspan citationid=\"CR90\" class=\"CitationRef\"\u003e2014\u003c/span\u003e, \u003cspan citationid=\"CR89\" class=\"CitationRef\"\u003e2015\u003c/span\u003e; Palmqvist et al., \u003cspan citationid=\"CR185\" class=\"CitationRef\"\u003e2022a\u003c/span\u003e). Figure\u0026nbsp;\u003cspan refid=\"Fig8\" class=\"InternalRef\"\u003e8\u003c/span\u003e also includes the mean estimates of carrying capacity for the Orce sites (3827.8 kg/km\u003csup\u003e2\u003c/sup\u003e for VM and 3307.6 kg/km\u003csup\u003e2\u003c/sup\u003e for BL/FN-3), obtained using an approach based on the Weibull model (see Rodr\u0026iacute;guez-G\u0026oacute;mez et al., \u003cspan citationid=\"CR204\" class=\"CitationRef\"\u003e2022\u003c/span\u003e, \u003cspan citationid=\"CR203\" class=\"CitationRef\"\u003e2024\u003c/span\u003e), and the estimates of annual rainfall, based on the composition of the herpetological assemblages in the case of BL/FN-3 (700 mm for BL and 738 mm for FN-3; Blain et al., \u003cspan citationid=\"CR32\" class=\"CitationRef\"\u003e2016\u003c/span\u003e) and on the abundance of nitrogen isotopes in the bone collagen retrieved from the fossils of large mammals from VM (800 mm; Palmqvist et al., \u003cspan citationid=\"CR187\" class=\"CitationRef\"\u003e2022c\u003c/span\u003e, \u003cspan citationid=\"CR184\" class=\"CitationRef\"\u003e2023\u003c/span\u003e). As deduced for the Baza Basin, the presence of thermal waters provided a variety of salts and trace elements. These mineral nutrients allowed the development of a diverse plant community and the translation of this biodiversity to the local trophic pyramid, thus configuring a sort of ecological \"Eden\" on a limited spatio-temporal scale. Given that some species of mammals, for example the hippopotamuses, used these thermal pools and watered them, their salinity (at least where the fossil remains are found) must have been of \u0026lt;\u0026thinsp;5 g/l (Palmqvist et al., \u003cspan citationid=\"CR185\" class=\"CitationRef\"\u003e2022a\u003c/span\u003e). In this environmental context, the ancient hominin settlers of the Baza Basin sought such places as preferential sources of water and a variety of trophic resources.\u003c/p\u003e \u003cp\u003e \u003c/p\u003e \u003cp\u003eFigure \u003cspan refid=\"Fig8\" class=\"InternalRef\"\u003e8\u003c/span\u003eB shows that although the productivity of the African National Parks and Game Reserves mainly depends on annual precipitation, as noted above, it also relates to soil type and to the presence of thermal waters, which contribute to increase the productivity of the terrestrial ecosystems and their herbivore biomass. Specifically, the areas with volcanic and Rift sediments have more positive residuals in the regression equation of Fig.\u0026nbsp;\u003cspan refid=\"Fig8\" class=\"InternalRef\"\u003e8\u003c/span\u003eA than those with other types of sediments (i.e., basement, floodplain, granite, Kalahari sands, and marine sediments; see also Bell, \u003cspan citationid=\"CR29\" class=\"CitationRef\"\u003e1981\u003c/span\u003e). Similarly, the areas with active hot springs show greater values of prey biomass than those in which thermal activity is absent (Fig.\u0026nbsp;\u003cspan refid=\"Fig8\" class=\"InternalRef\"\u003e8\u003c/span\u003eB).\u003c/p\u003e \u003c/div\u003e \u003cdiv id=\"Sec16\" class=\"Section2\"\u003e \u003ch2\u003e4.4 Correlation of the proposed model for the Baza Basin with other World sites\u003c/h2\u003e \u003cp\u003eA large-scale geographical position of 103 African, West Asian and European sites, which range in age from 7.5 to 0.35 Ma, shows a close relationship with those areas that show high tectonic and geothermal activity (Fig.\u0026nbsp;\u003cspan refid=\"Fig9\" class=\"InternalRef\"\u003e9\u003c/span\u003e). Most of these sites appear in areas that evidence intense geological activity related to seismotectonic, volcanic and hydrothermal phenomena during the Plio-Pleistocene, and even up to present days. These sites usually show tuffaceous levels, volcanic ashes, seismites, or lacustrine deposits potentially fed by thermal waters. This is also the case of the archaeological deposits of Java (Indonesia), where volcanism and seismicity are common and particularly intense (Delinom, \u003cspan citationid=\"CR61\" class=\"CitationRef\"\u003e2009\u003c/span\u003e).\u003c/p\u003e \u003cp\u003e \u003c/p\u003e \u003cp\u003eThese sites range in age from the appearance of the first hominins in Africa to the arrival of \u003cem\u003eHomo sapiens\u003c/em\u003e in Morocco, as detected in the Jebel Irhoud site (Hublin et al., \u003cspan citationid=\"CR125\" class=\"CitationRef\"\u003e2017\u003c/span\u003e). Figure\u0026nbsp;\u003cspan refid=\"Fig9\" class=\"InternalRef\"\u003e9\u003c/span\u003e shows that 88% of the sites are found in areas where volcanic, seismic, tectonic, and hydrothermal phenomena have been common since Late Miocene times (and in many cases up to present days). Of all these sites, 71% appear in sedimentary facies of lacustrine or fluvio-lacustrine origin, as happens throughout the East African Rift, while the other sites are found in karst cavities (21%) and in sedimentary formations of fluvial or marine-littoral origin (8%). Most paleoanthropological sites placed in geologically stable areas are found in karst cavities and fluvial paleo-terraces. If we consider the sedimentary origin and the tectonic environment of these sites, 67% of them are found in lacustrine or fluvio-lacustrine facies located in tectonically active areas with hot springs.\u003c/p\u003e \u003cp\u003eThe network of spatio-temporal nodes of hydrothermal phenomena in the Baza Basin seems to be related with the presence of hominin settlements, as evidenced by four archaeological localities: the late Early Pleistocene sites of Barranco Le\u0026oacute;n and Fuente Nueva-3, the Early-Middle Pleistocene site of Hu\u0026eacute;scar-1, and the early Middle Pleistocene site of C\u0026uacute;llar-Baza-1. This possible association has led us to the proposal of a new idea on the distribution of paleoanthropological sites on a global scale during the Plio-Pleistocene.\u003c/p\u003e \u003cp\u003eMany African and Eurasian sites appear in areas that evidence intense geological activity related to seismotectonic, volcanic and hydrothermal phenomena during the Plio-Pleistocene. In most cases, these areas remain tectonically active today (Fig.\u0026nbsp;\u003cspan refid=\"Fig9\" class=\"InternalRef\"\u003e9\u003c/span\u003e). Such scenarios can be compared with those that characterized the Plio-Pleistocene environments of the Baza Basin. Throughout the East African Rift Valley, from Mozambique-Malawi to Ethiopia and Eritrea, following the Red Sea and the Levantine Corridor to Anatolia and Georgia, the main archaeological deposits are found in volcanic contexts and scenarios characterized by intense tectonic activity, which resulted in geothermal anomalies during the Neogene-Quaternary (Morley et al., \u003cspan citationid=\"CR170\" class=\"CitationRef\"\u003e1992\u003c/span\u003e; Schl\u0026uuml;ter, \u003cspan citationid=\"CR227\" class=\"CitationRef\"\u003e1997\u003c/span\u003e; Adamia, \u003cspan citationid=\"CR3\" class=\"CitationRef\"\u003e2013\u003c/span\u003e; Glerum et al., \u003cspan citationid=\"CR104\" class=\"CitationRef\"\u003e2020\u003c/span\u003e). More specifically, a relationship between hydrothermal activity and paleoanthropological deposits of 1.7 Ma has been evidenced by microbial markers in Olduvai Gorge by Sistiaga et al. (\u003cspan citationid=\"CR240\" class=\"CitationRef\"\u003e2020\u003c/span\u003e), who used a similar model to the one proposed here for the Baza Basin. Such correspondence also occurs in several European sites linked to recent hydrothermal and seismo-tectonic activity, for example the Acheulean site of Rodafnidia, Lesvos Island (Galanidou et al., \u003cspan citationid=\"CR81\" class=\"CitationRef\"\u003e2013\u003c/span\u003e, \u003cspan citationid=\"CR82\" class=\"CitationRef\"\u003e2016\u003c/span\u003e). This shows the common relationship between areas with intense geothermal activity and the presence of archaeological sites with hominin remains and fossils of large mammals.\u003c/p\u003e \u003cp\u003eTectonic activity is a source of geomorphological variability (i.e., landscape roughness) and ecological diversity (Coppens, \u003cspan citationid=\"CR52\" class=\"CitationRef\"\u003e1994\u003c/span\u003e; Foley, \u003cspan citationid=\"CR74\" class=\"CitationRef\"\u003e1987\u003c/span\u003e; Gamble, \u003cspan citationid=\"CR83\" class=\"CitationRef\"\u003e1993\u003c/span\u003e) and has been linked to the first hominins that evolved in the African Rift, because it allowed them to obtain stable resources: the tectonic and volcanic movements that generated the East African Rift System resulted in a broken, hilly rough country of complex topography where the early hominins could hunt and hide (Bailey et al., \u003cspan citationid=\"CR23\" class=\"CitationRef\"\u003e2000\u003c/span\u003e, \u003cspan citationid=\"CR24\" class=\"CitationRef\"\u003e2011\u003c/span\u003e). Moreover, environmental roughness probably characterized the whole route of early hominin dispersal (King and Bailey, \u003cspan citationid=\"CR131\" class=\"CitationRef\"\u003e2006\u003c/span\u003e). We suggest here that tectonic and hydrothermal activity are significantly linked to the evolution of the human lineage due to the consumption strategies of hominins and their risk of exposure to confrontation with carnivores. In fact, carnivores could have predated the own hominins (Treves and Palmqvist, \u003cspan citationid=\"CR252\" class=\"CitationRef\"\u003e2007\u003c/span\u003e) and were also competitors in the access to meat, although they were also providers of carrion (Espigares et al., \u003cspan citationid=\"CR70\" class=\"CitationRef\"\u003e2013\u003c/span\u003e; Palmqvist et al., \u003cspan citationid=\"CR185\" class=\"CitationRef\"\u003e2022a\u003c/span\u003e, \u003cspan citationid=\"CR184\" class=\"CitationRef\"\u003e2023\u003c/span\u003e). According to the evidence unearthed from the fossil sites of Orce, the hominins developed a scavenging behavior as a successful strategy to obtain animal resources in the Baza Basin (Espigares et al., \u003cspan citationid=\"CR69\" class=\"CitationRef\"\u003e2019\u003c/span\u003e; Palmqvist et al., \u003cspan citationid=\"CR184\" class=\"CitationRef\"\u003e2023\u003c/span\u003e). Analyses on resource partitioning among the members of the carnivore guild suggest that scavenging was the optimal strategy for these hominin populations (Rodr\u0026iacute;guez-G\u0026oacute;mez et al., \u003cspan citationid=\"CR206\" class=\"CitationRef\"\u003e2016\u003c/span\u003e). Anatomical evidence from late Early Pleistocene sites younger than those of Orce, such as level TD6 from Gran Dolina in Sierra de Atapuerca, suggests that these humans had lesser throwing skills than modern humans, which would hinder a hunting activity based on throwing cobbles against prey, although it would not have limited other hunting strategies (Berm\u0026uacute;dez de Castro et al., 2020; however, for an opposite interpretation see Garc\u0026iacute;a-Mart\u0026iacute;nez et al., \u003cspan citationid=\"CR91\" class=\"CitationRef\"\u003e2021\u003c/span\u003e).\u003c/p\u003e \u003cp\u003eA heterogeneous environment would be useful for exploiting different ecological niches in the case of an opportunistic species like \u003cem\u003eHomo\u003c/em\u003e sp., relatively weak for a direct confrontation with the large carnivores. Moreover, these mosaic habitats would also provide protection from cursor predators in the open environments and from ambushing carnivores in the closed ones, as the hominins could inhabit the boundary zones between different environments, both for protecting themselves from predators and for exploiting the resources of these environments (see Treves and Palmqvist, \u003cspan citationid=\"CR252\" class=\"CitationRef\"\u003e2007\u003c/span\u003e). In ecology, the transition zones between different ecosystems are called ecotones. We propose that rather than a generalist species, the early members of \u003cem\u003eHomo\u003c/em\u003e would be specialized in exploiting the ecotones, thus showing adaptations to consume resources from different environments. According to an analysis of avifauna in Lower, Middle and Upper Paleolithic sites (Finlayson et al., \u003cspan citationid=\"CR72\" class=\"CitationRef\"\u003e2011\u003c/span\u003e), there is a significant association between hominin presence and mosaic habitats (i.e., habitats with many ecotones).\u003c/p\u003e \u003cp\u003eOn the other hand, hominin absence of Sierra de Atapuerca during the early Middle Pleistocene has been linked to a greater environmental homogeneity than during previous periods (Rodr\u0026iacute;guez et al., \u003cspan citationid=\"CR202\" class=\"CitationRef\"\u003e2011\u003c/span\u003e; Rodr\u0026iacute;guez-G\u0026oacute;mez et al., \u003cspan citationid=\"CR207\" class=\"CitationRef\"\u003e2014\u003c/span\u003e). Today, the areas with highest ecosystem diversity are two regions with active tectonics, the African Rift and the Himalayan range, in accordance with the ecoregions defined by Olson et al. (\u003cspan citationid=\"CR174\" class=\"CitationRef\"\u003e2001\u003c/span\u003e). The richness of ecoregions per unit area observed in the African Rift is probably lower nowadays than during the Plio-Pleistocene, because tectonic activity was higher at that time (Bailey et al., \u003cspan citationid=\"CR23\" class=\"CitationRef\"\u003e2000\u003c/span\u003e; King and Bailey, \u003cspan citationid=\"CR131\" class=\"CitationRef\"\u003e2006\u003c/span\u003e). This diversity of ecoregions seems to relate to the diversity in the values of abiotic and biotic variables that can be observed in the region. For these reasons, the African Rift would provide a high diversity of habitats in which a generalist or ecotone-specialist species, such as early \u003cem\u003eHomo\u003c/em\u003e, could have efficiently exploited these environments for resources and shelter. This model can be proposed for the ancient hominin populations found in the archaeological sites of the Baza Basin (for a review on the paleodemography of these populations, see Palmqvist et al., \u003cspan citationid=\"CR185\" class=\"CitationRef\"\u003e2022a\u003c/span\u003e).\u003c/p\u003e \u003cp\u003eFrom a paleoecological point of view, the Ngorongoro Crater in the African Rift Valley is a model that can be correlated with the sites of the Baza Basin and other deposits found in geologically active areas. In Ngorongoro, the hydrothermal influence of the crater lakes has a seasonal pattern. As a result, the populations of large mammals are dispersed in the wet season, while during the dry season they concentrate near the pastures that surround the hot springs. These waters lose their ecological influence at ~\u0026thinsp;700 m from their point of emergence, changing to alkaline (pH\u0026thinsp;=\u0026thinsp;~\u0026thinsp;8) waters with high saline contents (DeoCampo, \u003cspan citationid=\"CR62\" class=\"CitationRef\"\u003e2002\u003c/span\u003e). The list of species that live in the crater within a radius of ~\u0026thinsp;100 m around the thermal springs includes buffaloes, gazelles, hippopotamuses, rhinoceroses, zebras, elephants, antelopes, wild boars, hyenas, foxes, cheetahs, jackals, and lions. This fauna is strikingly similar to the one that inhabited the Baza Basin during the Plio-Pleistocene in terms of their distribution among modes of locomotion, feeding types and body size classes (Mendoza et al., \u003cspan citationid=\"CR165\" class=\"CitationRef\"\u003e2005\u003c/span\u003e).\u003c/p\u003e \u003c/div\u003e"},{"header":"5 Conclusions","content":"\u003cp\u003eHot springs have been important for faunal and hominin survival during the Plio-Pleistocene, favoring the dispersal of mammalian species as documented in the Baza Basin, which preserves one of the continental sequences with greater stratigraphic completeness of Europe, including the earliest archaeological evidence of hominin presence in the northern Mediterranean latitudes during the late Early Pleistocene. In this basin, hot springs form a network of spatio-temporal nodes, which are particularly concentrated in the Orce sub-basin, showing permanent siliceous waters with temperatures\u0026thinsp;\u0026gt;\u0026thinsp;35\u0026ordm; C. These hot springs played an important ecological role during the summer, when seasonal rain-fed springs became dry, and also helped to maintain locally warm temperatures in winter, thus aiding the survival of water dependent vertebrates adapted to warm and moist climates, including the members of early \u003cem\u003eHomo.\u003c/em\u003e\u003c/p\u003e \u003cp\u003eNearly one hundred Plio-Pleistocene archaeological and paleontological localities are recorded in the perimeter of the Baza Basin. Most of these sites were generated in connection with tectono-sedimentary processes, as evidenced by the presence of paleo-seismites, ancient hot springs and faulting. The lithological, mineralogical and geochemical composition of the chert levels present in the Orce sub-basin suggests a high-temperature hydrothermal origin for these deposits. Other types of sedimentary facies present in the basin, such as travertines, dark lutites or calcilutites, were influenced by this network of spatio-temporal nodes of hot springs, showing diverse geochemical and mineralogical indicators associated with this hydrothermal origin. The mineral nutrients provided by the hot springs prompted the development of a luxurious vegetation, which in turn allowed the development of a diverse community of large mammals, including the first hominins that dispersed in Western Europe.\u003c/p\u003e"},{"header":"Declarations","content":"\u003cp\u003e\u003cstrong\u003eCompliance with Ethical Standards\u003c/strong\u003e\u003c/p\u003e\n\u003cp\u003e\u003cstrong\u003eDisclosure of potential conflicts of interest:\u003c/strong\u003e On behalf of all authors, the corresponding author states that there is no conflict of interest.\u003c/p\u003e\n\u003cp\u003e\u003cstrong\u003eResearch involving Human Participants and/or Animals:\u003c/strong\u003e This research involved no human participants and/or animals.\u003c/p\u003e\n\u003cp\u003e\u003cstrong\u003eInformed consent:\u003c/strong\u003e All authors participated in this research, contributed to elaborating the manuscript contents and agreed in submitting it for publication in Journal of Iberian Geology.\u003c/p\u003e"},{"header":"References","content":"\u003col\u003e\u003cli\u003e\u003cspan\u003eAbbate, E., Albianelli, A., Azzaroli, A., Benvenuti, M., Tesfamariam, B., Bruni, P., et al. 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These sediments were deposited in fluvio-lacustrine environments and preserve huge assemblages of large mammals, including the oldest evidence of hominin presence in Western Europe at the late Early Pleistocene sites of Barranco Le\u0026oacute;n and Fuente Nueva-3, dated to \u0026sim;1.4 Ma. Since latest Miocene times, the basin was subject to intense tectonic activity, showing sedimentary deposits originated from ancient hot springs. These hot springs were developed in a spatio-temporal network related to tectonic fractures, showing a higher concentration in the Orce sub-basin during Late Pliocene to Holocene times. Compositional analyses of cherts and other facies associated with these hot springs show geochemical and mineralogical markers linked to hydrothermal scenarios. The contribution of hot springs in the Baza Basin resulted in rich and productive biotopes, which supported a high diversity and biomass of terrestrial mammals. Similar geological, hydrochemical and ecological environments are found along hotspot lines in East Africa, the Levantine Corridor and the circum-Mediterranean realm. We hypothesize here that the presence of these hot springs corridors favored the dispersal of hominins and other large mammals during the Plio-Pleistocene, which led to the first hominin arrival in Europe.\u003c/p\u003e","manuscriptTitle":"Plio-Pleistocene hydrothermal events of the Baza Basin (Betic Cordillera, SE Spain) and their paleoecological implications","msid":"","msnumber":"","nonDraftVersions":[{"code":1,"date":"2024-04-04 07:59:31","doi":"10.21203/rs.3.rs-4151726/v1","editorialEvents":[{"type":"communityComments","content":0},{"type":"decision","content":"Minor revisions","date":"2024-04-29T01:54:04+00:00","index":"","fulltext":""},{"type":"reviewerAgreed","content":"","date":"2024-04-01T16:08:07+00:00","index":0,"fulltext":""},{"type":"reviewersInvited","content":"","date":"2024-04-01T15:01:17+00:00","index":"","fulltext":""},{"type":"editorInvited","content":"Journal of Iberian Geology","date":"2024-03-29T13:31:57+00:00","index":"","fulltext":""},{"type":"editorAssigned","content":"","date":"2024-03-25T04:02:17+00:00","index":"","fulltext":""},{"type":"submitted","content":"Journal of Iberian Geology","date":"2024-03-23T04:27:35+00:00","index":"","fulltext":""}],"status":"published","journal":{"display":true,"email":"[email protected]","identity":"journal-of-iberian-geology","isNatureJournal":false,"hasQc":true,"allowDirectSubmit":false,"externalIdentity":"jibg","sideBox":"Learn more about [Journal of Iberian Geology](http://link.springer.com/journal/41513)","snPcode":"41513","submissionUrl":"https://www.editorialmanager.com/jibg/default2.aspx","title":"Journal of Iberian Geology","twitterHandle":"","acdcEnabled":true,"dfaEnabled":true,"editorialSystem":"em","reportingPortfolio":"Springer Hybrid","inReviewEnabled":true,"inReviewRevisionsEnabled":false}}],"origin":"","ownerIdentity":"9724deb8-2000-4412-93df-1f8373910dbc","owner":[],"postedDate":"April 4th, 2024","published":true,"recentEditorialEvents":[],"rejectedJournal":[],"revision":"","amendment":"","status":"under-review","subjectAreas":[],"tags":[],"updatedAt":"2024-07-08T09:22:21+00:00","versionOfRecord":[],"versionCreatedAt":"2024-04-04 07:59:31","video":"","vorDoi":"","vorDoiUrl":"","workflowStages":[]},"version":"v1","identity":"rs-4151726","journalConfig":"researchsquare"},"__N_SSP":true},"page":"/article/[identity]/[[...version]]","query":{"redirect":"/article/rs-4151726","identity":"rs-4151726","version":["v1"]},"buildId":"8U1c8b4HqxoKbykW_rLl7","isFallback":false,"isExperimentalCompile":false,"dynamicIds":[84888],"gssp":true,"scriptLoader":[]}