Silvopastoral Systems in Practice: Evidence from a Dairy Value Chain Intervention in Caquetá, Colombia | Research Square window.SnipcartSettings = { analytics: { enabled: false } }; (function() { var accessVector = localStorage.getItem('access_vector') || ''; window.dataLayer = window.dataLayer || []; if (accessVector) { window.dataLayer.push({ user: { profile: { profileInfo: { snid: accessVector } } } }); } })(); (function(w,d,s,l,i){w[l]=w[l]||[];w[l].push({'gtm.start':new Date().getTime(),event:'gtm.js'});var f=d.getElementsByTagName(s)[0],j=d.createElement(s),dl=l!='dataLayer'?'&l='+l:'';j.async=true;j.src='https://www.googletagmanager.com/gtm.js?id='+i+dl;f.parentNode.insertBefore(j,f);})(window,document,'script','dataLayer','GTM-K279D39R'); Browse Preprints In Review Journals COVID-19 Preprints AJE Video Bytes Research Tools Research Promotion AJE Professional Editing AJE Rubriq About Preprint Platform In Review Editorial Policies Our Team Advisory Board Help Center Sign In Submit a Preprint Cite Share Download PDF Article Silvopastoral Systems in Practice: Evidence from a Dairy Value Chain Intervention in Caquetá, Colombia Alexander Buritica This is a preprint; it has not been peer reviewed by a journal. https://doi.org/ 10.21203/rs.3.rs-8057588/v1 This work is licensed under a CC BY 4.0 License Status: Under Review Version 1 posted 10 You are reading this latest preprint version Abstract Food systems must raise output while reducing environmental harm, with livestock at the center of this challenge. In Colombia, cattle ranching dominates land use and is closely linked to deforestation, land degradation, and biodiversity loss, particularly in Amazonian frontier regions. This study evaluates a sustainability-oriented intervention in a Colombian dairy value chain that promoted silvopastoral systems (SPS) by bundling targeted inputs for paddock division and pasture improvement with technical assistance on rotational grazing and tree establishment. Using a differences-in-differences design combined with satellite imagery, panel data from supplier farms, and milk-quality records from processing plants, I estimate the impact of this intervention on SPS adoption and associated environmental and productive outcomes. Relative to control properties, treated farms reduced deforestation and exhibited increases in bird diversity and plant-biomass proxies. Support for paddock division accelerated rotational grazing and tree integration, improved perceived herd nutrition, and lowered disease incidence. Processing plants simultaneously recorded higher levels of protein, fat, and total solids in milk, providing downstream evidence consistent with improved feeding conditions, although not identified causally. The results indicate that integrated value chain support can deliver early gains and help align cattle production with conservation goals. JEL Classification Codes: O13, Q12, Q15, Q57. Biological sciences/Ecology Earth and environmental sciences/Ecology Earth and environmental sciences/Environmental sciences Livestock Market access Difference-in-differences Technology adoption Silvopastoral systems Deforestation Full Text Additional Declarations No competing interests reported. Supplementary Files draftSupplementaryMaterialsv2.docx Cite Share Download PDF Status: Under Review Version 1 posted Reviewers agreed at journal 07 Apr, 2026 Reviewers agreed at journal 02 Apr, 2026 Reviewers agreed at journal 06 Feb, 2026 Reviews received at journal 09 Jan, 2026 Reviewers agreed at journal 21 Dec, 2025 Reviewers invited by journal 09 Dec, 2025 Editor assigned by journal 09 Dec, 2025 Editor invited by journal 24 Nov, 2025 Submission checks completed at journal 20 Nov, 2025 First submitted to journal 20 Nov, 2025 You are reading this latest preprint version Research Square lets you share your work early, gain feedback from the community, and start making changes to your manuscript prior to peer review in a journal. As a division of Research Square Company, we’re committed to making research communication faster, fairer, and more useful. We do this by developing innovative software and high quality services for the global research community. Our growing team is made up of researchers and industry professionals working together to solve the most critical problems facing scientific publishing. Also discoverable on Platform About Our Team In Review Editorial Policies Advisory Board Help Center Resources Author Services Accessibility API Access RSS feed Manage Cookie Preferences © Research Square 2026 | ISSN 2693-5015 (online) Privacy Policy Terms of Service Do Not Sell My Personal Information {"props":{"pageProps":{"initialData":{"identity":"rs-8057588","acceptedTermsAndConditions":true,"allowDirectSubmit":false,"archivedVersions":[],"articleType":"Article","associatedPublications":[],"authors":[{"id":558112035,"identity":"97c35132-d8be-4e37-950c-cfe305d7866b","order_by":0,"name":"Alexander Buritica","email":"data:image/png;base64,iVBORw0KGgoAAAANSUhEUgAAAZAAAAAyAQMAAABI0h/eAAAABlBMVEX///8AAABVwtN+AAAACXBIWXMAAA7EAAAOxAGVKw4bAAAA3klEQVRIiWNgGAWjYFACHiA2AJHMB0AMorUYAEm2BFK0gJXyEKeegb+B95h0RcEfGf72Mx8/fimwy2dg7338Ap8WiQN8aZJngA6TOJO7WVrGINmygee4mQVeaw7wmEk2ALUYMORuY5YwYDZgkEhjw+tEebgW/jfPgFrqCWsxgGuRyGFj/GBwGKSF+QE+LYaH+YDONzDmkbjxzFiaweC4ARvPMTa8XpE73nvwZsMfOXv+/uSHH3/8qTbgZ29j/oBXDzMyGxRHQCvYJPBqQQaMP6Ba8dsyCkbBKBgFIw0AAEnxOu3kUfgkAAAAAElFTkSuQmCC","orcid":"","institution":"Alliance of Bioversity International and CIAT (CGIAR)","correspondingAuthor":true,"prefix":"","firstName":"Alexander","middleName":"","lastName":"Buritica","suffix":""}],"badges":[],"createdAt":"2025-11-07 13:38:08","currentVersionCode":1,"declarations":"","doi":"10.21203/rs.3.rs-8057588/v1","doiUrl":"https://doi.org/10.21203/rs.3.rs-8057588/v1","draftVersion":[],"editorialEvents":[],"editorialNote":"","failedWorkflow":false,"files":[{"id":98432944,"identity":"ecc9d0d9-efcc-430d-b32c-cd11773cfc28","added_by":"auto","created_at":"2025-12-17 16:50:07","extension":"docx","order_by":0,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":1404301,"visible":true,"origin":"","legend":"","description":"","filename":"draft20251119.docx","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/99302099e5bb57532ced4000.docx"},{"id":98433201,"identity":"5e3224c1-4471-4597-a14e-5c093d6827d9","added_by":"auto","created_at":"2025-12-17 16:50:26","extension":"json","order_by":1,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":3735,"visible":true,"origin":"","legend":"","description":"","filename":"af4661a7dc0e4b5498458d21d371c32e.json","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/04b1d5fc1b15b4be0246b751.json"},{"id":98212116,"identity":"2101b19a-509a-4450-9731-bb098000dd1c","added_by":"auto","created_at":"2025-12-15 09:44:12","extension":"docx","order_by":2,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":27019,"visible":true,"origin":"","legend":"","description":"","filename":"draftSupplementaryMaterialsv2.docx","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/2fb227188904400b7423876e.docx"},{"id":98433208,"identity":"5645e767-a931-4540-8ffa-e1330d82e408","added_by":"auto","created_at":"2025-12-17 16:50:27","extension":"xml","order_by":3,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":193758,"visible":true,"origin":"","legend":"","description":"","filename":"af4661a7dc0e4b5498458d21d371c32e1enriched.xml","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/cc52b6a7477798c6971203ff.xml"},{"id":98212107,"identity":"1ad7a340-720e-46cb-83a0-2227243598ad","added_by":"auto","created_at":"2025-12-15 09:44:11","extension":"png","order_by":4,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":151196,"visible":true,"origin":"","legend":"","description":"","filename":"floatimage1.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/796c7812547df1ad42e18d16.png"},{"id":98431275,"identity":"b0476f6e-351b-4494-a692-01e15ef2cc2e","added_by":"auto","created_at":"2025-12-17 16:47:25","extension":"png","order_by":5,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":27998,"visible":true,"origin":"","legend":"","description":"","filename":"floatimage10.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/2680917c67bf9ed774b80574.png"},{"id":98432347,"identity":"685f13e0-1d01-44d4-a620-938b2a080618","added_by":"auto","created_at":"2025-12-17 16:49:26","extension":"png","order_by":6,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":117345,"visible":true,"origin":"","legend":"","description":"","filename":"floatimage2.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/200f71cb20b645dafba5f2e0.png"},{"id":98212111,"identity":"8688eb74-0281-4ab8-b75c-f476039bf9bf","added_by":"auto","created_at":"2025-12-15 09:44:12","extension":"png","order_by":7,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":133251,"visible":true,"origin":"","legend":"","description":"","filename":"floatimage3.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/4917742b172541b5f3e15fe2.png"},{"id":98431374,"identity":"f89cd556-0641-4b81-a80b-c2aa0fedd428","added_by":"auto","created_at":"2025-12-17 16:47:36","extension":"jpeg","order_by":8,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":202351,"visible":true,"origin":"","legend":"","description":"","filename":"floatimage4.jpeg","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/f861b4b9aad68a12f1183679.jpeg"},{"id":98433612,"identity":"6ba94e61-3da5-4453-a84e-a835f5afec65","added_by":"auto","created_at":"2025-12-17 16:50:56","extension":"png","order_by":9,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":54926,"visible":true,"origin":"","legend":"","description":"","filename":"floatimage5.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/758524d22d2acf130ebfb64b.png"},{"id":98432841,"identity":"bfb047c6-5317-44a0-8b2f-ac5f28886725","added_by":"auto","created_at":"2025-12-17 16:50:02","extension":"png","order_by":10,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":392033,"visible":true,"origin":"","legend":"","description":"","filename":"floatimage6.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/7128720f46ef2c6df698e8c9.png"},{"id":98431646,"identity":"6463097e-9d9a-429f-ade3-dee2d7a18605","added_by":"auto","created_at":"2025-12-17 16:48:05","extension":"png","order_by":11,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":68356,"visible":true,"origin":"","legend":"","description":"","filename":"floatimage7.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/f7ccc17ebd47a85b6561cf96.png"},{"id":98432367,"identity":"59004e02-3400-4899-ab93-f55f8b9fae33","added_by":"auto","created_at":"2025-12-17 16:49:28","extension":"png","order_by":12,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":49449,"visible":true,"origin":"","legend":"","description":"","filename":"floatimage8.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/24542635bf1bfb718dc00d1d.png"},{"id":98431406,"identity":"dae1b3e3-4b08-43d7-9f87-19270a4c748b","added_by":"auto","created_at":"2025-12-17 16:47:38","extension":"png","order_by":13,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":80784,"visible":true,"origin":"","legend":"","description":"","filename":"floatimage9.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/f72fedef5468e6279d5d3589.png"},{"id":98432703,"identity":"c2c233d9-93f3-4218-8217-fcbb4d052dc7","added_by":"auto","created_at":"2025-12-17 16:49:49","extension":"jpeg","order_by":14,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":285,"visible":true,"origin":"","legend":"","description":"","filename":"groupimage1.jpeg","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/e9562d7f0542756274a93894.jpeg"},{"id":98432330,"identity":"c8f70404-6d98-4aa1-a56b-c6e5a73a3fb6","added_by":"auto","created_at":"2025-12-17 16:49:24","extension":"png","order_by":15,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":43518,"visible":true,"origin":"","legend":"","description":"","filename":"Onlinefloatimage1.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/99e51647deabdfb823af7781.png"},{"id":98432820,"identity":"20dd544b-7e5a-43be-a602-c11d90f2f4c6","added_by":"auto","created_at":"2025-12-17 16:50:00","extension":"png","order_by":16,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":15046,"visible":true,"origin":"","legend":"","description":"","filename":"Onlinefloatimage10.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/00f817392d3810735fe38053.png"},{"id":98212128,"identity":"63d639c6-09a5-466c-bba1-4b04adfb21bf","added_by":"auto","created_at":"2025-12-15 09:44:12","extension":"png","order_by":17,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":61529,"visible":true,"origin":"","legend":"","description":"","filename":"Onlinefloatimage2.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/ff7e7be8f99936cbe9bf6049.png"},{"id":98431616,"identity":"b30bba43-901e-49a9-b361-78c00a6f21e0","added_by":"auto","created_at":"2025-12-17 16:48:02","extension":"png","order_by":18,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":69163,"visible":true,"origin":"","legend":"","description":"","filename":"Onlinefloatimage3.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/ababf1db8083fdb7d0ba38ab.png"},{"id":98212121,"identity":"e8ca9a1a-5c24-4e6d-9fab-e62cbd772405","added_by":"auto","created_at":"2025-12-15 09:44:12","extension":"png","order_by":19,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":34381,"visible":true,"origin":"","legend":"","description":"","filename":"Onlinefloatimage4.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/530ebff91dd5c3f219d759f3.png"},{"id":98212127,"identity":"78bff9ac-4c5f-4d81-8183-72c751985b23","added_by":"auto","created_at":"2025-12-15 09:44:12","extension":"png","order_by":20,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":18782,"visible":true,"origin":"","legend":"","description":"","filename":"Onlinefloatimage5.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/6e88a083433fa5a13b1bd2c1.png"},{"id":98432735,"identity":"c10fc7c9-6e00-4f1e-bad9-7793a7175dd3","added_by":"auto","created_at":"2025-12-17 16:49:54","extension":"png","order_by":21,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":79357,"visible":true,"origin":"","legend":"","description":"","filename":"Onlinefloatimage6.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/19bd7e7236069d2b00fd9f8c.png"},{"id":98212124,"identity":"9b617e78-695b-4728-bc99-6d525d13f328","added_by":"auto","created_at":"2025-12-15 09:44:12","extension":"png","order_by":22,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":17901,"visible":true,"origin":"","legend":"","description":"","filename":"Onlinefloatimage7.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/bda9c7d9da7b0080e9dbf519.png"},{"id":98212130,"identity":"d620257a-fcb3-4efa-a773-16ea50bd5241","added_by":"auto","created_at":"2025-12-15 09:44:12","extension":"png","order_by":23,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":14357,"visible":true,"origin":"","legend":"","description":"","filename":"Onlinefloatimage8.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/650e632584c206db6cec03ee.png"},{"id":98212126,"identity":"6a3ae2b3-42a3-404e-b1ef-62567a716040","added_by":"auto","created_at":"2025-12-15 09:44:12","extension":"png","order_by":24,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":23389,"visible":true,"origin":"","legend":"","description":"","filename":"Onlinefloatimage9.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/4761ea49f2a1e71193d2a415.png"},{"id":98432334,"identity":"5cdbb56e-c812-4fcf-a9a5-937aff8b2295","added_by":"auto","created_at":"2025-12-17 16:49:25","extension":"png","order_by":25,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":317,"visible":true,"origin":"","legend":"","description":"","filename":"Onlinegroupimage1.png","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/39004f93c21392258df91c52.png"},{"id":98432342,"identity":"67227c44-7259-40c0-948a-dad7363bf71f","added_by":"auto","created_at":"2025-12-17 16:49:26","extension":"xml","order_by":26,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":187883,"visible":true,"origin":"","legend":"","description":"","filename":"af4661a7dc0e4b5498458d21d371c32e1structuring.xml","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/b3e943e9293cc4bbda28d253.xml"},{"id":98212136,"identity":"a6c17302-4af9-4652-bbab-ec1717ea1e42","added_by":"auto","created_at":"2025-12-15 09:44:12","extension":"html","order_by":27,"title":"","display":"","copyAsset":false,"role":"acdc-reference","size":208793,"visible":true,"origin":"","legend":"","description":"","filename":"earlyproof.html","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/dcd7929a67742327e2fece43.html"},{"id":98622302,"identity":"efeb56e3-2674-4255-a862-ec53a96e7ebe","added_by":"auto","created_at":"2025-12-19 16:51:22","extension":"pdf","order_by":1,"title":"","display":"","copyAsset":false,"role":"manuscript-pdf","size":1299361,"visible":true,"origin":"","legend":"","description":"","filename":"draft20251119.pdf","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1_covered_03622695-30a1-410f-9f06-2ac7b84b7430.pdf"},{"id":98432245,"identity":"55e07976-b53b-4577-9b82-dcf0801c9f07","added_by":"auto","created_at":"2025-12-17 16:49:16","extension":"docx","order_by":0,"title":"","display":"","copyAsset":false,"role":"supplement","size":27019,"visible":true,"origin":"","legend":"","description":"","filename":"draftSupplementaryMaterialsv2.docx","url":"https://assets-eu.researchsquare.com/files/rs-8057588/v1/f674b2669ee492914eadece4.docx"}],"financialInterests":"No competing interests reported.","formattedTitle":"Silvopastoral Systems in Practice: Evidence from a Dairy Value Chain Intervention in Caquetá, Colombia","fulltext":[],"fulltextSource":"","fullText":"","funders":[],"hasAdminPriorityOnWorkflow":false,"hasManuscriptDocX":false,"hasOptedInToPreprint":true,"hasPassedJournalQc":"","hasAnyPriority":false,"hideJournal":false,"highlight":"","institution":"","isAcceptedByJournal":false,"isAuthorSuppliedPdf":true,"isDeskRejected":"","isHiddenFromSearch":false,"isInQc":false,"isInWorkflow":false,"isPdf":true,"isPdfUpToDate":true,"isWithdrawnOrRetracted":false,"journal":{"display":true,"email":"
[email protected]","identity":"scientific-reports","isNatureJournal":false,"hasQc":true,"allowDirectSubmit":false,"externalIdentity":"scirep","sideBox":"Learn more about [Scientific Reports](http://www.nature.com/srep/)","snPcode":"","submissionUrl":"","title":"Scientific Reports","twitterHandle":"","acdcEnabled":true,"dfaEnabled":true,"editorialSystem":"stoa","reportingPortfolio":"Scientific Reports","inReviewEnabled":true,"inReviewRevisionsEnabled":true},"keywords":"Livestock, Market access, Difference-in-differences, Technology adoption, Silvopastoral systems, Deforestation","lastPublishedDoi":"10.21203/rs.3.rs-8057588/v1","lastPublishedDoiUrl":"https://doi.org/10.21203/rs.3.rs-8057588/v1","license":{"name":"CC BY 4.0","url":"https://creativecommons.org/licenses/by/4.0/"},"manuscriptAbstract":"\u003cp\u003eFood systems must raise output while reducing environmental harm, with livestock at the center of this challenge. In Colombia, cattle ranching dominates land use and is closely linked to deforestation, land degradation, and biodiversity loss, particularly in Amazonian frontier regions. This study evaluates a sustainability-oriented intervention in a Colombian dairy value chain that promoted silvopastoral systems (SPS) by bundling targeted inputs for paddock division and pasture improvement with technical assistance on rotational grazing and tree establishment. Using a differences-in-differences design combined with satellite imagery, panel data from supplier farms, and milk-quality records from processing plants, I estimate the impact of this intervention on SPS adoption and associated environmental and productive outcomes. Relative to control properties, treated farms reduced deforestation and exhibited increases in bird diversity and plant-biomass proxies. Support for paddock division accelerated rotational grazing and tree integration, improved perceived herd nutrition, and lowered disease incidence. Processing plants simultaneously recorded higher levels of protein, fat, and total solids in milk, providing downstream evidence consistent with improved feeding conditions, although not identified causally. The results indicate that integrated value chain support can deliver early gains and help align cattle production with conservation goals.\u003c/p\u003e\n\u003cp\u003e\u003cstrong\u003eJEL Classification Codes:\u003c/strong\u003e O13, Q12, Q15, Q57.\u003c/p\u003e","manuscriptTitle":"Silvopastoral Systems in Practice: Evidence from a Dairy Value Chain Intervention in Caquetá, Colombia","msid":"","msnumber":"","nonDraftVersions":[{"code":1,"date":"2025-12-15 09:44:07","doi":"10.21203/rs.3.rs-8057588/v1","editorialEvents":[{"type":"communityComments","content":0},{"type":"reviewerAgreed","content":"259442831390926343727337952074171933128","date":"2026-04-07T13:02:44+00:00","index":"hide","fulltext":""},{"type":"reviewerAgreed","content":"236596487126651543772512458849974211976","date":"2026-04-02T13:02:11+00:00","index":"hide","fulltext":""},{"type":"reviewerAgreed","content":"36288726162003314102685266903910806277","date":"2026-02-06T16:02:09+00:00","index":"hide","fulltext":""},{"type":"editorInvitedReview","content":"","date":"2026-01-09T19:53:20+00:00","index":"hide","fulltext":""},{"type":"reviewerAgreed","content":"13864393060085303127878026649277559445","date":"2025-12-21T15:39:59+00:00","index":"hide","fulltext":""},{"type":"reviewersInvited","content":"","date":"2025-12-09T13:58:37+00:00","index":"","fulltext":""},{"type":"editorAssigned","content":"","date":"2025-12-09T13:57:10+00:00","index":"","fulltext":""},{"type":"editorInvited","content":"","date":"2025-11-24T13:22:23+00:00","index":"","fulltext":""},{"type":"checksComplete","content":"","date":"2025-11-20T09:23:58+00:00","index":"","fulltext":""},{"type":"submitted","content":"Scientific Reports","date":"2025-11-20T09:20:06+00:00","index":"","fulltext":""}],"status":"published","journal":{"display":true,"email":"
[email protected]","identity":"scientific-reports","isNatureJournal":false,"hasQc":true,"allowDirectSubmit":false,"externalIdentity":"scirep","sideBox":"Learn more about [Scientific Reports](http://www.nature.com/srep/)","snPcode":"","submissionUrl":"","title":"Scientific Reports","twitterHandle":"","acdcEnabled":true,"dfaEnabled":true,"editorialSystem":"stoa","reportingPortfolio":"Scientific Reports","inReviewEnabled":true,"inReviewRevisionsEnabled":true}}],"origin":"","ownerIdentity":"6de206e9-b542-476f-b2ec-46e9266305de","owner":[],"postedDate":"December 15th, 2025","published":true,"recentEditorialEvents":[],"rejectedJournal":[],"revision":"","amendment":"","status":"under-review","subjectAreas":[{"id":59406919,"name":"Biological sciences/Ecology"},{"id":59406920,"name":"Earth and environmental sciences/Ecology"},{"id":59406921,"name":"Earth and environmental sciences/Environmental sciences"}],"tags":[],"updatedAt":"2025-12-15T09:44:07+00:00","versionOfRecord":[],"versionCreatedAt":"2025-12-15 09:44:07","video":"","vorDoi":"","vorDoiUrl":"","workflowStages":[]},"version":"v1","identity":"rs-8057588","journalConfig":"researchsquare"},"__N_SSP":true},"page":"/article/[identity]/[[...version]]","query":{"redirect":"/article/rs-8057588","identity":"rs-8057588","version":["v1"]},"buildId":"8U1c8b4HqxoKbykW_rLl7","isFallback":false,"isExperimentalCompile":false,"dynamicIds":[84888],"gssp":true,"scriptLoader":[]}
Text is read by the "Ask this paper" AI Q&A widget below.
Extraction quality varies by source — PMC NXML preserves structure
cleanly, OA-HTML may include some navigation residue, and OA-PDF can
have broken hyphenation. The publisher copy
(via DOI)
is the canonical version.