How Large Cooperative Groups Avoid Local Competition

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Abstract

Large cooperative groups are a common sight in nature. Their existence is puzzling, however, because local competition should keep groups relatively small. A simple but untested way large groups can avoid local competition is by increasing their resource base. We conducted a systematic review and phylogenetic meta-analysis to look for evidence of this effect in wild populations of cooperatively breeding birds. Across 634 groups from 29 species, group size and resource availability (territory area) were strongly positively correlated (Pearson’s r = 0.52). Furthermore, when a specific group changed size, its territory changed size correspondingly (N = 34 groups). Our results support the prediction that large groups can avoid local competition by increasing their resource base and explain a sizeable fraction of group size variation in nature (R2 = 0.27).
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This is a Preprint and has not been peer reviewed. This is version 1 of this Preprint. You must log in to post a comment. There are no comments or no comments have been made public for this article. This is a Preprint and has not been peer reviewed. This is version 1 of this Preprint. Add a Comment You must log in to post a comment. Comments There are no comments or no comments have been made public for this article. Large cooperative groups are a common sight in nature. Their existence is puzzling, however, because local competition should keep groups relatively small. A simple but untested way large groups can avoid local competition is by increasing their resource base. We conducted a systematic review and phylogenetic meta-analysis to look for evidence of this effect in wild populations of cooperatively breeding birds. Across 634 groups from 29 species, group size and resource availability (territory area) were strongly positively correlated (Pearson’s r = 0.52). Furthermore, when a specific group changed size, its territory changed size correspondingly (N = 34 groups). Our results support the prediction that large groups can avoid local competition by increasing their resource base and explain a sizeable fraction of group size variation in nature (R2 = 0.27). https://doi.org/10.32942/X2QM1Q Behavior and Ethology, Ecology and Evolutionary Biology, Evolution Comparative study, Aves, helper, alloparent, social evolution Published: 2026-01-19 20:30 Last Updated: 2026-01-19 20:30 CC BY Attribution 4.0 International Conflict of interest statement: We declare no conflict of interest Data and Code Availability Statement: Data, code currently available here: https://github.com/pipdowning/Birds_Local_Competition/tree/main Language: English

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last seen: 2026-05-20T01:45:00.602351+00:00