Scaling diversity, abundance and energy: The Equilibrium Theory of Biodiversity Dynamics

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This chapter introduces the Equilibrium Theory of Biodiversity Dynamics (ETBD), a framework for understanding large-scale biodiversity patterns by examining how speciation and extinction rates depend on population size, diversity, and environmental factors.

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Abstract

This preprint of a book chapter presents the newly proposed Equilibrium Theory of Biodiversity Dynamics (ETBD), whose aim is to conceptualize large-scale dynamics of species richness via addressing the population size-dependence of speciation and extinction rates, the resulting diversity-dependence of these rates, and their modulation by the environment. It provides the most general framework for understanding large-scale biodiversity patterns such as the latitudinal diversity gradient (LDG) and temporal patterns of biodiversity changes. The theory has been published elsewhere in its full form that includes all the derivations (Okie & Storch 2004, American Naturalist https://doi.org/10.1086/733103 ), but here it is presented in a simpler and user-friendly way, focusing on its major implications comprising macroecological scaling relationships between energy (or resource) availability, species richness and community abundance.

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last seen: 2026-05-20T01:45:00.602351+00:00