Mapping the next forest generation – the potential of national forest inventory data for identifying regeneration gaps

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Abstract

In light of global change and forest disturbances, there is an increasing recognition of the importance of forest regeneration to ensure future generations of trees. However, despite the importance of forest regeneration, there is a lack in spatial information on the current availability of trees in the seedling and sapling stage. In this study, we aimed to evaluate the potential to predict species-specific forest regeneration densities using regeneration data typically recorded within National Forest Inventories (NFIs). We then calculated three indicators for regeneration quantity and quality to locate potential gaps of regeneration under a changing climate. We successfully calibrated regeneration density models for 22 tree species using generalised additive models (GAMs) using regeneration density data from the 2012 German NFI and 44 environmental predictors. Subsequently, the models were used to create regeneration density maps for the German forest area at high spatial resolution (1 ha). Regeneration gaps were evaluated in terms of low total density (less than 1,000 ha-1), low species richness (≤2 species) and a high proportion (≥75%) of regeneration at high future cultivation risk. Our results indicate gaps in terms of total regeneration density and species richness for 13.4% and 47.1% of the forest area of Germany, respectively. A lack of climate-adapted species was found for 25.2%, exemplarily assessed for the Bavarian forest area. Along this example, we show how such results can be used to identify areas that require additional silvicultural intervention in order to increase the resilience of future forests. Our study highlights the potential of NFI data, particularly that on forest regeneration, and demonstrates the applicability of regeneration indicator maps for forest management and policymakers in times of change.
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Abstract

- In face of global change and increasing forest disturbances, forest regeneration is crucial for ensuring future generations of trees and resilient forest ecosystems. However, spatially explicit information on the current availability and climate suitability of seedlings and saplings remains scarce. - We assessed the potential to predict species-specific forest regeneration densities at high spatial resolution (1 ha) by calibrating generalized additive models (GAMs) using regeneration data from the German National Forest Inventory (NFI) and 44 environmental predictors. Regional regeneration gaps were then identified based on three indicators: low total density (<1,000 ha-1), low species richness (≤2 species) and a high proportion (≥75%) of regeneration at high future cultivation risk. - For 22 tree species, we obtained regeneration density models that performed well in spatially blocked cross-validation. We were therefore able to generate regeneration density and indicator maps for a major part of the tree species. - The indicator maps revealed considerable regeneration gaps. 13.4% of Germany’s forest area has low regeneration density, 47.1% has low species richness, and 25.2% of the Bavarian forest area lacks climate-adapted regeneration. - Our study demonstrates the potential of NFI regeneration data and its applicability for monitoring forest regeneration over large spatial scales. The regeneration indicator maps show that silvicultural interventions should prioritise increasing tree species richness and the proportion of species adapted to climate change. However, as regeneration gaps vary from region to region, management and policy must be adapted accordingly to ensure future forest resilience. - Synthesis and applications: Our study provides the first nationwide, high-resolution assessment of forest regeneration, offering a valuable baseline for monitoring forest development. The regeneration density and indicator maps enable forest managers and policymakers to identify regeneration deficits, prioritise adaptive management interventions, and contribute to the development of climate-resilient forests. DOI https://doi.org/10.32942/X2GS8X Subjects Forest Biology, Forest Management, Forest Sciences, Other Forestry and Forest Sciences, Plant Sciences

Keywords

forest regeneration, species distribution models SDMs, generalized additive models GAMs, sapling density, species richness, climate-adapted species, cultivation risk Dates Published: 2025-05-31 09:33 Last Updated: 2026-03-30 07:34 Older Versions License CC-BY Attribution-NonCommercial 4.0 International Additional Metadata Conflict of interest statement: None Data and Code Availability Statement: All code supporting the findings of this study is openly available on Zenodo (https://doi.org/10.5281/zenodo.15552196) and GitHub (https://github.com/LeonieCG/GermanRegenerationMaps2012). The data used to calibrate our species-specific regeneration models was compiled from the German national forest inventory as well as several metadata sources, originally collected by various other institutions. As far as we were permitted, we have republished the data on Zenodo (https://doi.org/10.5281/zenodo.15550864) and provided the code to run the models with the reduced set of environmental variables. Language: English

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License: CC-BY-NC-4.0